The Influence of Grazing on Soil Multifunctionality of the Rehabilitated Grasslands on the Loess Plateau

doi:10.11733/j.issn.1007-0435.2025.12.024

Acta Agrestia Sinica ›› 2025, Vol. 33 ›› Issue (12): 4119-4128.DOI: 10.11733/j.issn.1007-0435.2025.12.024

• 2025-12-28 • Previous Articles

The Influence of Grazing on Soil Multifunctionality of the Rehabilitated Grasslands on the Loess Plateau

MING Jiao^1,3, ZHANG Bing-chang^1,2, WANG Guo-hua¹, ZHAO Yun-ge^3,4

1. Shanxi Normal University, College of Geographical Sciences, Taiyuan, Shanxi Province 030031, China;
2. Research Center of Ecological Environment of Middle Reaches of Yellow River, Geography Science College, Shanxi Normal University, Taiyuan, Shanxi Province 030031, China;
3. State Key Laboratory of Soil and Water Conservation and Desertification Control, Institute of Soil and Water Conservation, College of Soil and Water Conservation Science and Engineering (Institute of Soil and Water Conservation), Northwest A&F University, Yangling, Shannxi Province 712100, China;
4. Institute of Soil and Water Conservation, Chinese Academy of Sciences and Ministry of Water Resources, Yangling, Shannxi Province 712100, China

Received:2025-05-22 Revised:2025-07-17 Published:2025-12-01

放牧对黄土高原退耕草地土壤多功能性的影响

明姣^1,3, 张丙昌^1,2, 王国华¹, 赵允格^3,4

1. 山西师范大学地理科学学院, 山西太原 030031;
2. 山西师范大学黄河中游生态环境研究中心, 山西太原 030031;
3. 西北农林科技大学水土保持科学与工程学院(水土保持研究所)水土保持与荒漠化整治全国重点实验室, 陕西杨凌 712100;
4. 中国科学院水利部水土保持研究所, 陕西杨凌 712100

通讯作者: 赵允格，E-mail：zyunge@ms.iswc.ac.cn
作者简介:明姣（1987-），女，汉族，陕西西安人，博士，讲师，主要从事干旱半干旱生态系统土壤生态过程及调控研究，E-mail：mingjiao@sxnu.edu.cn；
基金资助:
国家自然科学基金项目（42377357；41830758）；水土保持与荒漠化整治全国重点实验室开放基金 Z2010025001-KJ2517资助

Abstract

Abstract: Soil multifunctionality （SMF） has a crucial impact on grassland ecosystem services and regional sustainable development. This study explored the effects of different grazing intensities （G₁-G₅： 2.2， 3.0， 4.2， 6.7， 16.7 sheep·hm^-2） and grazing durations on the SMF of the rehabilitated grassland on the Loess Plateau through fencing grazing experiments. The results showed that light grazing （2.2-3.0 sheep·hm^-2） could promote the soil organic carbon （SOC） and total nitrogen （TN） accumulation， and presented a time accumulation effect. Light grazing increased the content of ammonium nitrogen （NH₄⁺-N）， but had no significant effect on nitrate nitrogen （NO₃^--N） and available phosphorus （AP）. Meanwhile， light grazing significantly increased soil C∶P and N∶P. C∶P and N∶P increased by 8.03%-9.81% and 10.87%-11.59% respectively under G₁-G_2，while under G_5， C∶P and N∶P significantly reduced. With the increase of grazing duration， both C∶P and N∶P under G₁-G₅ grazing decreased， indicating that phosphorus might become the limiting factor of soil nutrient supply in this area. Under each grazing intensity， the SMF of the surface soil did not change significantly within two years， and the soil showed strong system stability. In conclusion， moderate grazing （≤3.0 sheep·hm^-2） can increase soil nutrients and maintain the stability of SMF in the short term， which provides a scientific basis for the scientific management and moderate utilization of grassland converted from degraded farmland on the Loess Plateau.

Key words: Grazing, Soil multifunctionality, Cropland conversion to grassland, Loess Plateau, Post-Grain for Green era

摘要： 土壤多功能性（Soil multifunctionality， SMF）对草地生态系统服务和区域可持续发展具有关键影响。本研究通过围栏放牧试验探究了不同放牧强度（G₁-G₅：2.2，3.0，4.2，6.7，16.7羊·hm^-2）及放牧时间对黄土高原退耕草地SMF的影响。结果表明，轻度放牧（2.2~3.0 羊·hm^-2）可促进土壤有机碳（Soil organic carbon， SOC）和全氮（Total nitrogen，TN）积累，并呈现时间累积效应；轻度放牧增加铵态氮（NH₄⁺-N）含量，而对硝态氮（NO₃^--N）及速效磷（Available phosphorus，AP）含量无显著影响；同时，轻度放牧显著提高土壤C∶P和N∶P，G₁-G₂放牧处理C∶P增加8.03%~9.81%，N∶P增加10.87%~11.59%，G₅放牧处理显著降低C∶P和N∶P；随着放牧历时增加，G₁-G₅放牧处理C∶P和N∶P均降低，表明磷可能成为该区土壤养分供应的限制因子。各放牧强度下表层土壤SMF在两年内未发生显著变化，土壤显示出较强的系统稳定性。综上，适度放牧（≤3.0 羊·hm^-2）可在短期内增加土壤养分，维持SMF稳定，这为黄土高原退耕草地科学管理及适度利用提供科学依据。

关键词: 放牧, 土壤多功能性, 退耕草地, 黄土高原, 后退耕时代

CLC Number:

S812.2

MING Jiao, ZHANG Bing-chang, WANG Guo-hua, ZHAO Yun-ge. The Influence of Grazing on Soil Multifunctionality of the Rehabilitated Grasslands on the Loess Plateau[J]. Acta Agrestia Sinica, 2025, 33(12): 4119-4128.

明姣, 张丙昌, 王国华, 赵允格. 放牧对黄土高原退耕草地土壤多功能性的影响[J]. 草地学报, 2025, 33(12): 4119-4128.

References

[1] DENG L，SHANGGUAN Z P. High quality developmental approach for soil and water conservation and ecological protection on the Loess Plateau[J]. Frontiers of Agricultural Science and Engineering，2021，8（4）：501
[2] LIU G B，SHANGGUAN Z P，YAO W Y，et al. Ecological effects of soil conservation in Loess Plateau[J]. Bulletin of Chinese Academy of Sciences，2017，32（1）：11-19 刘国彬，上官周平，姚文艺，等. 黄土高原生态工程的生态成效[J]. 中国科学院院刊，2017，32（1）：11-19
[3] CHEN Y P，WANG K B，LIN Y S，et al. Balancing green and grain trade[J]. Nature Geoscience，2015，8（10）：739-741
[4] SHAO M A，JIA X X，WANG Y Q，et al. A review of studies on dried soil layers in the Loess Plateau[J]. Advances in Earth Science，2016，31（1）：14-22 邵明安，贾小旭，王云强，等. 黄土高原土壤干层研究进展与展望[J]. 地球科学进展，2016，31（1）：14-22
[5] FENG X M，FU B J，PIAO S L，et al. Revegetation in China’s Loess Plateau is approaching sustainable water resource limits[J]. Nature Climate Change，2016，6（11）：1019-1022
[6] LI X，XIAO Q F，NIU J Z，et al. Process-based rainfall interception by small trees in Northern China：the effect of rainfall traits and crown structure characteristics[J]. Agricultural and Forest Meteorology，2016，218：65-73
[7] BÜNEMANN E K，BONGIORNO G，BAI Z G，et al. Soil quality-A critical review[J]. Soil Biology and Biochemistry，2018，120：105-125
[8] LEI F Y，LI X S，TAO Y，et al. Characterization of soil multifunctionality and its determining factors under moss crust cover in the arid regions of Northwest China[J]. Arid Zone Research，2024，41（5）：812-820 雷菲亚，李小双，陶冶，等. 西北干旱区藓类结皮覆盖下土壤多功能性特征及影响因子[J]. 干旱区研究，2024，41（5）：812-820
[9] GOU Y N，NAN Z B. The impacts of grazing on the soil microorganism population of grassland[J]. Acta Prataculturae Sinica，2015，24（ 10）：194-205 苟燕妮，南志标. 放牧对草地土壤微生物的影响[J]. 草业学报，2015，24（10）：194-205
[10] FıRıNCıOĞLU H K，SEEFELDT S S，ŞAHIN B. The effects of long-term grazing exclosures on range plants in the central Anatolian Region of Turkey[J]. Environmental Management，2007，39（3）：326-337
[11] RITCHIE M E. Plant compensation to grazing and soil carbon dynamics in a tropical grassland[J]. PeerJ，2014，2：e233
[12] WU J H，PANG H，ZHUO Y，et al. Effects of vegetation degradation on carbon and nitrogen distribution of roots-soil system in temperate typical steppe[J].Acta Ecologica Sinica，2018，38（15）：5340-5350 邬嘉华，庞惠，卓义，等. 植被退化对温带典型草原根系-土壤系统碳氮分配的影响[J]. 生态学报，2018，38（15）：5340-5350
[13] HICKMAN K R，HARTNETT D C. Effects of grazing intensity on growth，reproduction，and abundance of three palatable forbs in Kansas tallgrass prairie[J]. Plant Ecology，2002，159（1）：23-33
[14] LYU P，ZUO X A，ZHANG J，et al. Effects of grazing intensity on vegetation in sandy grassland of horqin[J]. Journal of Desert Research，2016，36（1）：34-39 吕朋，左小安，张婧，等. 放牧强度对科尔沁沙地沙质草地植被的影响[J]. 中国沙漠，2016，36（1）：34-39
[15] YAN R R，WEI Z J，XIN X P， et al. The effect of different grazing systems on soil nutrient contents in dessert steppe[J].Acta Ecologica Sinica，2010，30（1）：43-51 闫瑞瑞，卫智军，辛晓平，等. 放牧制度对荒漠草原生态系统土壤养分状况的影响[J]. 生态学报，2010，30（1）：43-51
[16] CHEN L L，WANG K X，BAOYIN T. Effects of grazing and mowing on vertical distribution of soil nutrients and their stoichiometry （C∶N∶P） in a semi-arid grassland of North China[J]. Catena，2021，206：105507
[17] HE F C，DONG S K，YANG J J，et al. Rotational grazing can enhance the soil available nutrients rather than plant diversity to promote ANPP of alpine steppe[J]. Agriculture，Ecosystems and Environment，2025，387：109617
[18] LAI L M，KUMAR S. A global meta-analysis of livestock grazing impacts on soil properties[J]. PLoS One，2020，15（8）：e0236638
[19] ZHANG Y X，DU Y L，LIU X Q，et al. Impact of grazing activities on grassland ecosystems in Inner Mongolia[J]. Acta Ecologica Sinica，2025，45（10）：4963-4974 张雨欣，杜宜霖，刘小茜，等. 放牧活动对内蒙古草地生态系统的影响[J]. 生态学报，2025，45（10）：4963-4974
[20] LI W，MA X X，MA N，et al. Effects of grazing intensity on stoichiomerty of biological soil crusts in the hilly Loess Plateau Region[J]. Acta Agrestia Sinica，2021，29（11）：2547-2555 李雯，马昕昕，马宁，等. 放牧强度对黄土丘陵区生物结皮土壤化学计量学特征的影响[J]. 草地学报，2021，29（11）：2547-2555
[21] YAN Y Z，ZHANG Q，BUYANTUEV A，et al. Plant functional β diversity is an important mediator of effects of aridity on soil multifunctionality[J]. Science of the Total Environment，2020，726：138529
[22] SANDERSON M A，SKINNER R H，BARKER D J，et al. Plant species diversity and management of temperate forage and grazing land ecosystems[J]. Crop Science，2004，44（4）：1132-1144
[23] MAESTRE F T，QUERO J L，GOTELLI N J，et al. Plant species richness and ecosystem multifunctionality in global drylands[J]. Science，2012，335（6065）：214-218
[24] MING J，YANG G，ZHAO Y G，et al. Effects of grazing on soil water recharge of rehabilitated grassland under natural rainfall on the Loess Plateau，Northwest China[J]. Chinese Journal of Applied Ecology，2023，34（6）：1555-1562 明姣，杨光，赵允格，等. 放牧对降雨条件下黄土高原退耕草地土壤水分补给的影响[J]. 应用生态学报，2023，34（6）：1555-1562
[25] LI H Y，BAI J B，XUE Q. Present situation，existing problems and suggestions of grassland contracting in Yan’an City，Shaanxi Province[J]. Animal Husbandry and Feed Science，2011，32（7）：65-66 李海燕，白锦保，薛清. 陕西省延安市草原承包的现状及存在问题和建议[J]. 畜牧与饲料科学，2011，32（7）：65-66
[26] CHEN F R，CHENG J M，LIU W，et al. Effects of disturbances on organic soil carbon in the typical grassland of Loess Plateau[J]. Acta Agrestia Sinica，2012，20（2）：298-304，311 陈芙蓉，程积民，刘伟，等. 不同干扰对黄土高原典型草原土壤有机碳的影响[J]. 草地学报，2012，20（2）：298-304，311
[27] ZHAO L P，TAN S T，BAI X，et al. Effect of grazing and grazing exclusion management on offspring recruitment in a semiarid steppe on the Loess Plateau[J]. Journal of Northwest A & F University （Natural Science Edition），2016，44（9）：27-34 赵凌平，谭世图，白欣，等. 封育和放牧对黄土高原典型草原繁殖更新的影响[J]. 西北农林科技大学学报（自然科学版），2016，44（9）：27-34
[28] FENG H Z，CUI Z F，YANG M Y. Severd considerations on forage cultivation in the seven counties in northern Yan’an prefecture[J]. Pratacultural Science，2005，22（4）：63-65 冯宏珠，崔志峰，杨明耀. 延安北部七县牧草栽培的几点思考[J]. 草业科学，2005，22（4）：63-65
[29] MA N，ZHAO Y G，MA X X，et al. Response of soil carbon pool management index of the revegetated grasslands to grazing in the hilly Loess Plateau Region[J]. Acta Agrestia Sinica， 2022， 30（5）：1043-1053 马宁，赵允格，马昕昕，等. 黄土丘陵区退耕草地土壤碳库管理指数对放牧的响应[J]. 草地学报，2022，30（5）：1043-1053
[30] BAO S D. Soil Agrochemical Analysis[M]. 3rd Edition. Beijing： China Agriculture Press，2000：25-97 鲍士旦. 土壤农化分析[M]. 3版. 北京：中国农业出版社，2000：25-97
[31] WANG S S，ZHAO Y G，SHI Y F，et al. Impact of short-term grazing disturbance on nitrogen accumulation of biological soil crusts in the hilly Loess Plateau region，China[J]. Chinese Journal of Applied Ecology，2017，28（12）：3848-3854 王闪闪，赵允格，石亚芳，等. 短期放牧干扰对黄土丘陵区生物结皮土壤氮素累积的影响[J]. 应用生态学报，2017，28（12）：3848-3854
[32] GAO L Q，ZHAO Y G，XU M X，et al. The effects of biological soil crust succession on soil ecological stoichiometry characteristics[J]. Acta Ecologica Sinica，2018，38（2）：678-688 高丽倩，赵允格，许明祥，等. 生物土壤结皮演替对土壤生态化学计量特征的影响[J]. 生态学报，2018，38（ 2）：678-688
[33] SOLIVERES S，MAESTRE F T，ELDRIDGE D J，et al. Plant diversity and ecosystem multifunctionality peak at intermediate levels of woody cover in global drylands[J]. Global Ecology and Biogeography，2014，23（12）：1408-1416
[34] VALENCIA E，MAESTRE F T，LE BAGOUSSE-PINGUET Y，et al. Functional diversity enhances the resistance of ecosystem multifunctionality to aridity in Mediterranean drylands[J]. New Phytologist，2015，206（2）：660-671
[35] LI Z W，MIAO Y J，ZONG N. Effects of grazing exclusion on plant diversity and ecosystem multifunctionality of alpine grasslands in northern Xizang [J]. Acta Agrestia Sinica，2025，33（2）：596-608 李振威，缪雨珏，宗宁. 围栏封育对藏北高寒草地植物多样性与生态系统多功能性的影响 [J].草地学报，2025，33（2）：596-608
[36] BYRNES J E K，GAMFELDT L，ISBELL F，et al. Investigating the relationship between biodiversity and ecosystem multifunctionality：challenges and solutions [J]. Methods in Ecology and Evolution，2014，5（2）：111-124
[37] WANG L，DELGADO-BAQUERIZO M，WANG D L，et al. Diversifying livestock promotes multidiversity and multifunctionality in managed grasslands[J]. Proceedings of the National Academy of Sciences，2019，116（13）：6187-6192
[38] LIU Y Z，ZHAO X Q，DONG Q M，et al. Research progress on the effects of grazing on grassland ecosystem structure and function[J]. Acta Agrestia Sinica，2023，31（8）：2253-2262 刘玉祯，赵新全，董全民，等. 放牧对草地生态系统结构与功能影响的研究进展[J]. 草地学报，2023，31（8）：2253-2262
[39] SUN H，ZHAO Y G，LIU G L，et al. Effects of grazing intensity on the grassland vegetation community characteristics in the hilly Loess Plateau Region[J]. Acta Agrestia Sinica，2022，30（4）：810-817 孙会，赵允格，刘广亮，等. 放牧强度对黄土丘陵区草地植物群落特征的影响[J]. 草地学报，2022，30（4）：810-817
[40] WU J X，ZHANG Y H，LI S Y，et al. Effects of long-term grazing on different fractions of organic phosphorus content in desert grassland in Inner Mongolia[J]. Acta Agrestia Sinica，2024，32（5）：1479-1488 吴佳芯，张育涵，李邵宇，等. 长期放牧对内蒙古荒漠草原土壤不同组分有机磷含量的影响[J]. 草地学报，2024，32（5）：1479-1488
[41] SU Z S，SUN Y F，FU J J，et al. Effects of grazing intensity on soil nutrient of Kobresia pygmaea meadow in Tibet Plateau[J]. Pratacultural Science，2015，32（3）：322-328 苏振声，孙永芳，付娟娟，等. 不同放牧强度下西藏高山嵩草草甸土壤养分的变化[J]. 草业科学，2015，32（3）：322-328
[42] LI T Q，CAO J R，LIU X N，et al. Response of soil enzyme stoichiometry to grazing and identification of soil limiting nutrients in typical steppe of Nei Mongol，China[J]. Chinese Journal of Plant Ecology，2025，49（1）：19-29 李天琦，曹继容，柳小妮，等. 内蒙古典型草原土壤酶化学计量与限制性养分对放牧的响应[J]. 植物生态学报，2025，49（1）：19-29
[43] SONG X H，WANG Y H，WANG Z W，et al. Effects of different grazing intensity on aboveground biomass and litterfall amount in Inner Mongolia Stipa breviflora desert steppe[J]. Grassland and Prataculture，2019，31（2）：52-58 宋晓辉，王悦骅，王占文，等. 不同放牧强度对短花针茅荒漠草原地上生物量和枯落物量的影响[J]. 草原与草业，2019，31（2）：52-58
[44] PAKEMAN R J，THWAITES R H，LE DUC M G，et al.. Vegetation re-establishment on land previously subject to control of Pteridium aquilinum by herbicide[J]. Applied Vegetation Science，2000，3（1）：95-104
[45] SCASTA J D，GERGENI T，MACZKO K，et al. Adaptive grazing and animal density implications for stocking rate and drought in northern mixed-grass prairie[J]. Livestock Science， 2023， 269：105184
[46] MING J. Soil water of the rehabilitated grasslands on the Loess Plateau and its response to grazing interference[D]. Beijing：University of Chinese Academy of Sciences （The Research Center of Soil and Water Conservation and Ecological Environment，Chinese Academy of Sciences and Ministry of Education），2023：21-33 明姣. 黄土高原退耕封禁草地土壤水分对放牧干扰的响应[D]. 北京：中国科学院大学（中国科学院教育部水土保持与生态环境研究中心），2023：21-33
[47] BARDGETT R D，MOMMER L，DE VRIES F T. Going underground：root traits as drivers of ecosystem processes[J]. Trends in Ecology and Evolution，2014，29（12）：692-699

The Influence of Grazing on Soil Multifunctionality of the Rehabilitated Grasslands on the Loess Plateau

放牧对黄土高原退耕草地土壤多功能性的影响

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