The Influence of Cervus elaphus kansuensis Grazing on Leaf-Soil Ecological Stoichiometry Characteristics of Plant Community in A Long-Term Enclosed Grassland of Ningxia

doi:10.11733/j.issn.1007-0435.2026.04.022

Acta Agrestia Sinica ›› 2026, Vol. 34 ›› Issue (4): 1391-1402.DOI: 10.11733/j.issn.1007-0435.2026.04.022

The Influence of Cervus elaphus kansuensis Grazing on Leaf-Soil Ecological Stoichiometry Characteristics of Plant Community in A Long-Term Enclosed Grassland of Ningxia

MA Xiao-rui¹, LI Wei², AN Ke³, ZHANG Meng-jie⁴, TIAN Hui-hui⁵, WANG Yi-bo⁵, TUO Hang-hang¹, YE Fa-ming^6,7, YIN Zi-ming¹, YANG Qing^6,7, HAN Xin-ning⁸, LI Wei¹

1. State Key Laboratory of Soil and Water Conservation and Desertification Control, College of Soil and Water Conservation Science and Engineering, Northwest A&F University, Yangling, Shaanxi Province 712100, China;
2. Ningxia Rencundu Revetment Forest Farm, Lingwu, Ningxia 751400, China;
3. Dingbian County Meteorological Bureau, Dingbian, Shaanxi Province 718600, China;
4. College of Agriculture, Ningxia University, Yinchuan, Ningxia 750021, China;
5. College of Grassland Science and Management, Northwest A&F University, Yangling, Shaanxi Province 712100, China;
6. Institute of Soil and Water Conservation, Chinese Academy of Sciences and Ministry of Water Resources, Yangling, Shaanxi Province 712100, China;
7. University of Chinese Academy of Sciences, Beijing 110494, China;
8. College of Resources, Environment and Life Sciences, Ningxia Normal University, Guyuan, Ningxia 756000, China

Received:2025-09-19 Revised:2025-11-17 Published:2026-04-15

马鹿放牧对云雾山封育草地植物-土壤系统化学计量特征的影响

马晓瑞¹, 李玮², 安可³, 张梦婕⁴, 田慧慧⁵, 王亦波⁵, 拓行行¹, 叶发明^6,7, 尹子鸣¹, 杨庆^6,7, 韩新宁⁸, 李伟¹

1. 西北农林科技大学水土保持科学与工程学院/水土保持与荒漠化整治全国重点实验室, 陕西杨凌 712100;
2. 宁夏仁存渡护岸林场, 宁夏灵武 751400;
3. 定边县气象局, 陕西定边 718600;
4. 宁夏大学农学院, 宁夏银川 750021;
5. 西北农林科技大学草业与草原学院, 陕西杨凌 712100;
6. 中国科学院水利部水土保持研究所, 陕西杨凌 712100;
7. 中国科学院大学, 北京 110494;
8. 宁夏师范大学资源环境与生命科学学院, 宁夏固原 756000

通讯作者: 韩新宁，E-mail：hanxinning@163.com；李伟，E-mail：liwei2013@nwsuaf.edu.cn
作者简介:马晓瑞（2001-），女，汉族，山东临沂人，硕士研究生，主要从事草地生态学等研究，E-mail：maxr@nwafu.edu.cn；
基金资助:
国家自然科学基金项目（42277464）；国家重点研发计划项目（2022YFF1302800）；陕西省重点研发项目（2024SF-YBXM-545）；宁夏自然科学基金（2025AAC030610）；2025榆林市科技计划项目（2025-CXY-045）资助

Abstract

Abstract: Investigating the effects of grazing on the ecological chemometric characteristics of carbon （C）， nitrogen （N）， and phosphorus （P） in the plant-soil continuum of long-term enclosed grasslands is crucial for sustainable grassland management. This study investigated a 38-year-old enclosed grassland in Yunwu Mountain， Ningxia， by systematically analyzing ecological-metabolic ratios and their coupling relationships at the community level （community-weighted average） for leaves and soil across four grazing intensities of Gansu Cervus canadensis （Cervus elaphus kansuensis）： ungrazed， light grazing， moderate grazing， and heavy grazing. Results indicated：（1） Moderate grazing most effectively promoted N and P accumulation. Plant growth shifted from P limitation under no grazing and combined N-P limitation under light/moderate grazing to N limitation under heavy grazing；（2） With increasing grazing intensity， both topsoil soil organic carbon （SOC） and C∶N ratio in the topsoil （0~10 cm） showed an initial increase followed by a decrease， peaking at moderate grazing levels at 40.40 g·kg^-1 and 12.08， respectively. Conversely， soil N， C∶P， and N∶P ratios were the lowest at light grazing levels at 3.22 g·kg^-1， 48.15， and 4.43， respectively. All soil parameters decreased with increasing soil depth. （3） Leaf C and N contents showed negative correlations with most soil SOC， N， P， and their ratios， while leaf P content was correlated positively with these parameters. Soil depth influenced both plant leaf and soil chemical parameters， whereas grazing intensity primarily affected soil-related indicators. In summary， grazing significantly influenced the chemical-nutrient characteristics of the plant-soil system in the long-term enclosed grassland of Yunwu Mountain. Moderate grazing promoted plant growth and soil nutrient accumulation， representing a critical grazing intensity threshold for maintaining regional grassland health. These findings provide data support and theoretical basis for further optimizing grazing management and promoting the sustainable use of long-term closed-pasture grasslands.

Key words: Cervus elaphus grazing, Steppe, Long-term fencing, Ecological stoichiometry

摘要： 探讨放牧对长期封育草地植物-土壤连续体碳（C）、氮（N）、磷（P）生态化学计量学特征的影响对草地可持续管理具有重要意义。本研究以宁夏云雾山封育38年的草地为研究对象，通过设置未放牧、轻度放牧、中度放牧、重度放牧4种甘肃马鹿（Cervus elaphus kansuensis）放牧强度，系统分析群落水平（即群落加权平均）叶片和土壤的生态化学计量比及其耦合关系。结果表明：（1）中度放牧最有利于N， P养分的积累，且植物生长的养分限制由未放牧的P限制和轻、中度放牧的N-P共同限制，转变为重度放牧的N限制；（2）随放牧强度的增加，表层（0~10 cm）土壤有机碳（Soil organic carbon，SOC）、C∶N均呈先升后降的趋势，并在中度放牧达峰值，分别为40.40 g·kg^-1，12.08，而土壤N， C∶P， N∶P则在轻度放牧最低，分别为3.22 g·kg^-1， 48.15， 4.43，随土层加深，所有土壤指标均呈递减趋势；（3）叶片C， N含量与多数土壤SOC， N， P及计量比呈负相关，而叶片P含量与这些指标则呈正相关；土层深度同时影响植物叶片和土壤化学计量，而放牧强度则主要作用于土壤相关指标。综上，马鹿放牧显著影响了云雾山长期封育草地植物-土壤系统的生态化学计量特征，中度放牧有利于植物生长和土壤养分积累，是维持区域草地健康的关键放牧强度阈值，这些结果为进一步优化放牧管理，促进长期封育草地的可持续利用提供了数据支撑和理论依据。

关键词: 马鹿放牧, 典型草原, 长期封育, 生态化学计量

CLC Number:

S812

MA Xiao-rui, LI Wei, AN Ke, ZHANG Meng-jie, TIAN Hui-hui, WANG Yi-bo, TUO Hang-hang, YE Fa-ming, YIN Zi-ming, YANG Qing, HAN Xin-ning, LI Wei. The Influence of Cervus elaphus kansuensis Grazing on Leaf-Soil Ecological Stoichiometry Characteristics of Plant Community in A Long-Term Enclosed Grassland of Ningxia[J]. Acta Agrestia Sinica, 2026, 34(4): 1391-1402.

马晓瑞, 李玮, 安可, 张梦婕, 田慧慧, 王亦波, 拓行行, 叶发明, 尹子鸣, 杨庆, 韩新宁, 李伟. 马鹿放牧对云雾山封育草地植物-土壤系统化学计量特征的影响[J]. 草地学报, 2026, 34(4): 1391-1402.

References

[1] HAO T X，YANG M，YU G R. Coordinating the functions of five reservoirs of grassland ecosystem and constructing national ecological security barrier[J]. Science Foundation in China，2023（4）：603-612 郝天象，杨萌，于贵瑞. 统筹草地生态系统五库功能构建国家生态安全屏障[J]. 中国科学基金， 2023（4）：603-612
[2] HASSAN N，WANG Z W. Paralleled grazing and mowing differentially affected plant community diversity and productivity in a semi-arid grassland[J]. Ecological Processes，2024，13（1）：62
[3] SUN C Z，TANG S M，WANG Z，et al. Effect of enclosure of temperate grassland on plant community[J]. Acta Agrestia Sinica，2023（5）：1481-1489 孙崇之，唐士明，王珍，等. 温带草原围封对植物群落的影响[J]. 草地学报，2023（5）：1481-1489
[4] CUEVAS J G，LE QUESNE C. Low vegetation recovery after short-term cattle exclusion on Robinson Crusoe Island[J]. Plant Ecology，2006，183（1）：105-124
[5] HUANG M. Effects of long-term fencing on vegetation， soil， and resilience of artificial grassland in the “black soil land” of the three-river headwater region[D]. Lanzhou：Lanzhou University，2021：1-3 黄梅. 三江源区“黑土滩”人工草地长期围栏封育对植被，土壤和恢复力的影响[D]. 兰州：兰州大学，2021：1-3
[6] HOBBS R J，HIGGS E，HARRIS J A. Novel ecosystems： implications for conservation and restoration[J]. Trends in Ecology & Evolution，2009，24（11）：599-605
[7] REN Q，AI Y，HU J，et al. Effects of different yak grazing intensities on soil and plant biomass in an alpine meadow on the Qinghai-Tibetan Plateau[J]. Acta Ecologica Sinica，2021，41（17）：6862-6870 任强，艾鷖，胡健，等. 不同强度牦牛放牧对青藏高原高寒草地土壤和植物生物量的影响[J]. 生态学报，2021，41（17）：6862-6870
[8] LIU P，MA H，ZHI Y B，et al. Ecological stoichiometric differences of nine typical eremophyte species[J]. Arid Zone Research，2018，35（1）：207-216 刘珮，马慧，智颖飙，等. 9种典型荒漠植物生态化学计量学特征分析[J]. 干旱区研究，2018，35（1）：207-216
[9] YONGFEI BAI J W. Grazing alters ecosystem functioning and C： N： P stoichiometry of grasslands along a regional precipitation gradient[J]. Journal of Applied Ecology，2012，49（6）：1204-1215
[10] VEEN G F，DE VRIES S，BAKKER E S，et al. Grazing-induced changes in plant–soil feedback alter plant biomass allocation[J]. Oikos，2014，123（7）：800-806
[11] CHEN H，ZHU Q A，PENG C H，et al. The impacts of climate change and human activities on biogeochemical cycles on the Qinghai-Tibetan Plateau[J]. Global Change Biology，2013，19（10）：2940-2955
[12] ZHOU C H. Effects of grazing intensity on leaf C， N， and P stoichiometric characteristics and nutrient resorption of alpine meadow plants[D]. Lanzhou：Lanzhou University，2023：2-4 周彩虹. 放牧强度对高寒草甸植物叶片C，N，P化学计量特征及养分重吸收的影响[D]. 兰州：兰州大学， 2023：2-4
[13] GÜSEWELL S. N： P ratios in terrestrial plants： variation and functional significance[J]. The New Phytologist，2004，164（2）：243-266
[14] SONG M J. Effects of grazing Gansu red deer， yak， and Tibetan sheep on Salix cupularis shrub vegetation and soil in the eastern Qilian Mountains[D]. Lanzhou： Gansu Agricultural University， 2021：2-5 宋美娟. 放牧甘肃马鹿与牦牛藏羊对东祁连山杯腺柳灌丛植被和土壤的影响[D]. 兰州：甘肃农业大学，2021：2-5
[15] LIU J. Response of plant diversity and productivity to fencing and management practices in the grasslands of the semi-arid Loess Region[D]. Yangling：Northwest A & F University，2020：13-14 刘建. 半干旱黄土区草地植物多样性和生产力对封育和管理措施的响应[D]. 杨凌：西北农林科技大学，2020：13-14
[16] LI M Y. Evaluation of the effects of fencing duration on soil infiltration capacity in the grasslands of Yunwu Mountain[D]. Yangling：Northwest A & F University，2022：2-8 李梦瑶. 云雾山草地封育年限对土壤入渗能力影响评价[D]. 杨凌：西北农林科技大学，2022：2-8
[17] TUO H H. Assessment of ecosystem multifunctionality in typical grasslands of the Loess Plateau under the disturbance of red deer grazing[D]. Yangling：Northwest A & F University，2024：10-11 拓行行. 马鹿放牧干扰下黄土高原典型草原生态系统多功能性评估[D]. 杨凌：西北农林科技大学，2024：10-11
[18] FANG J Y，WANG X P，SHEN Z H，et al. Methods and protocols for plant community inventory[J]. Biodiversity Science，2009，17（6）：533-548 方精云，王襄平，沈泽昊，等. 植物群落清查的主要内容、方法和技术规范[J]. 生物多样性，2009，17（6）：533-548
[19] ERIC GARNIER J C. Plant functional markers capture ecosystem properties during secondary succession[J]. Ecology，2004，85（9）：2630-2637
[20] HE M，ZHOU G Y，YUAN T F，et al. Grazing intensity significantly changes the C： N： P stoichiometry in grassland ecosystems[J]. Global Ecology and Biogeography，2020，29（2）：355-369
[21] YU R P，ZHANG W P，FORNARA D A，et al. Contrasting responses of nitrogen： phosphorus stoichiometry in plants and soils under grazing： A global meta-analysis[J]. Journal of Applied Ecology，2021，58（5）：964-975
[22] LIU Y Z. Impacts of grazing on carbon， nitrogen， and phosphorus stoichiometric characteristics in global grassland ecosystems： A meta-analysis[J]. Chinese Journal of Applied Ecology，2022，33（5）：1251-1259 刘玉祯，刘文亭，杨晓霞，等. 放牧对全球草地生态系统碳氮磷化学计量特征影响的Meta分析[J]. 应用生态学报，2022， 33（5）：1251-1259
[23] MILCHUNAS W K L D G. Quantitative effects of grazing on vegetation and soils over a global range of environments[J]. Ecological Monographs，1993，63（4）：327-366
[24] DONG X Y，FU H，LI X D，et al. Effects of grazing and enclosure on plant biomass and carbon，nitrogen and phosphorus storage in typical grassland of Loess Plateau[J]. Acta Prataculturae Sinica，2010（2）：175-182 董晓玉，傅华，李旭东，等. 放牧与围封对黄土高原典型草原植物生物量及其碳氮磷贮量的影响[J]. 草业学报，2010（2）：175-182
[25] ZHANG P J，HUANG J H，MU L，et al. Effect of nitrogen and water addition on productivity of desert grassland under grazing background[J]. Acta Ecologica Sinica，2022，42（13）：5458-5470 张璞进，黄建辉，木兰，等. 氮水添加对放牧背景下荒漠草原生产力的影响[J]. 生态学报，2022，42（13）：5458-5470
[26] WANG L，XU H K，ZHANG H，et al. Grazing and mowing affect the carbon-to-nitrogen ratio of plants by changing the soil available nitrogen content and soil moisture on the meadow steppe，China[J]. Plants，2022，11（3）：286-286
[27] CAO J，YAN R R，CHEN X Y，et al. Grazing affects the ecological stoichiometry of the plant–soil–microbe system on the Hulunber steppe，China[J]. Sustainability，2019，11（19）：5226-5226
[28] DU E Z，TERRER C，PELLEGRINI A F A，et al. Global patterns of terrestrial nitrogen and phosphorus limitation[J]. Nature Geoscience，2020，13（3）：221-226
[29] KOERSELMAN W，MEULEMAN A F M. The vegetation N∶P ratio： a new tool to detect the nature of nutrient limitation[J]. The Journal of Applied Ecology，1996，33（6）：1441
[30] WANG Y L，ZHU W Y，HOU J J，et al. Effects of grazing on leaf traits of four degradation indicator plants in Guinan County[J]. Pratacultural Science，2020，37（3）：423-431 王娅琳，朱文琰，侯将将，等.放牧对贵南县4种退化指示植物叶片性状的影响[J]. 草业科学，2020，37（3）：423-431
[31] MCSHERRY M E，RITCHIE M E. Effects of grazing on grassland soil carbon： a global review[J]. Global Change Biology，2013，19（5）：1347-1357
[32] WANG Z J，CHEN M，WANG H H，et al. Effects of grazing intensity on the stoichiometric ratio of carbon，nitrogen and phosphorus in the soil of Reaumuria soongorica shrub and its reproductive characteristics[J]. Journal of Desert Research，2025（2）：119-128 汪正蛟，陈敏，王怀海，等. 放牧强度对红砂（Reaumuria soongorica）灌丛土壤碳氮磷化学计量比及繁殖性状的影响[J]. 中国沙漠， 2025（2）：119-128
[33] LI S Q. Response of soil nutrient characteristics to grazing utilization in alpine meadows of the northeastern margin of the Qinghai-Tibetan Plateau[D]. Lanzhou：Lanzhou University，2014：18-26 李世卿. 青藏高原东北边缘地区高寒草甸土壤养分特征对放牧利用的响应[D]. 兰州：兰州大学，2014：18-26
[34] FANG X，GUO X L，ZHENG R B， et al. [J]. Research of Soil and Water Conservation，2020，27（2）：9-14 方昕，郭雪莲，郑荣波，等. 不同放牧干扰对滇西北高原泥炭沼泽土壤生态化学计量特征的影响[J]. 水土保持研究，2020，27（2）：9-14
[35] JIA T T，YUAN X X，ZHAO H，et al. Effects of grazing on dominant plants and soil nitrogen and phosphorus contents in alpine meadow[J]. Chinese Journal of Grassland，2013（6）：80-85 贾婷婷，袁晓霞，赵洪，等. 放牧对高寒草甸优势植物和土壤氮磷含量的影响[J]. 中国草地学报， 2013（6）：80-85
[36] LI H Y. Effects of different grazing regimes on soil organic carbon fractions and their stability in typical grasslands[D]. Hohhot：Inner Mongolia University，2022：31-35 李航宇. 不同放牧方式对典型草原土壤有机碳组分及其稳定性的影响[D]. 呼和浩特：内蒙古大学， 2022：31-35
[37] SA R，CAO F，AO T G，et al. The effects of short term grazing intensities on soil physical and chemical properties in the inner monglia typical steppe[J]. Journal of Inner Mongolia Agricultural University （Natural Science Edition），2014，35（5）：75-79 萨仁高娃，曹芙，敖特根，等. 短期放牧强度对典型草原土壤理化性质的影响[J]. 内蒙古农业大学学报（自然科学版），2014，35（5）：75-79
[38] ZHANG Z X，YU Y L，LI Y Q，et al. Effects of grazing intensity on soil organic carbon and its spatial heterogeneity in desert steppe of Inner Mongolia[J]. Acta Ecologica Sinica，2021（15）：6257-6266 张子胥，于倚龙，李永强，等. 放牧强度对内蒙古荒漠草原土壤有机碳及其空间异质性的影响[J]. 生态学报，2021（15）：6257-6266
[39] MA Z W，WANG H，YU Y W，et al. Short-term response of forage regeneration to mowing in winter pasture of Gansu red deer[J]. Pratacultural Science，2018（1）：157-164 马周文，王宏，于应文，等. 甘肃马鹿冬季牧场牧草再生性对刈割的短期响应[J]. 草业科学，2018（1）：157-164
[40] HU H J. Studies on the molecular ecology and nutritional ecology of Tibet red deer （Cervus wallichii）[D]. Harbin：Northeast Forestry University，2016：45-57 胡贺娇. 西藏马鹿（Cervus wallichii）分子生态学与营养生态学研究[D]. 哈尔滨：东北林业大学，2016：45-57
[41] LI P，LING T W，YANG Z Q，et al. Eco-stoichiometric characteristics of carbon，nitrogen and phosphorus in leaves-soil of Pinus massoniana plantations with different forest ages[J]. Journal of Southwest Forestry University（Natural Sciences），2023（1）：88-98 李鹏，零天旺，杨章旗，等. 不同林龄马尾松人工林叶片-土壤碳氮磷生态化学计量特征研究[J]. 西南林业大学学报（自然科学），2023（1）：88-98
[42] NIE M H，SHEN Y，LU Y，et al. Eco-stoichiometric characteristics of leaf-soil of dominant plants in different communities in desert grassland area of Yanchi County，Ningxia[J]. Acta Agrestia Sinica，2021（1）：131-140 聂明鹤，沈艳，陆颖，等. 宁夏盐池县荒漠草原区不同群落优势植物叶片-土壤生态化学计量特征[J]. 草地学报，2021（1）：131-140
[43] BA G D. Ecological stoichiometric characteristics of leaf-soil in tree， shrub， and herb layers of natural forests in Kanas[D]. Urumqi：Xinjiang University，2023：32-36 巴格登. 喀纳斯天然林乔灌草叶片-土壤生态化学计量特征研究[D]. 乌鲁木齐：新疆大学，2023：32-36
[44] BA G D，WANG W D，XU Z L，et al. Stoichiometric characteristics of carbon，nitrogen and phosphorus in leaves and soil of Arbor，shrub and grass in Kanas natural forest[J]. Acta Ecologica Sinica，2023（21）：8749-8758 巴格登，王文栋，许仲林，等. 喀纳斯天然林乔灌草叶片及土壤碳氮磷化学计量特征[J]. 生态学报，2023（21）：8749-8758
[45] CHEN C，ZHANG S J，LI L D，et al. Stoichiometric characteristics of carbon，nitrogen and phosphorus in plant leaves，litter and soil during vegetation restoration in the middle subtropical zone[J]. Chinese Journal of Plant Ecology，2019（8）：658-671 陈婵，张仕吉，李雷达，等. 中亚热带植被恢复阶段植物叶片、凋落物、土壤碳氮磷化学计量特征[J]. 植物生态学报，2019（8）：658-671
[46] NACRY P，BOUGUYON E，GOJON A. Nitrogen acquisition by roots： physiological and developmental mechanisms ensuring plant adaptation to a fluctuating resource[J]. Plant and Soil，2013，370（1）：1-29
[47] ZHANG C W，WANG X，AN K，et al. Ecological stoichiometric characteristics of leaf-soil and differences in plant homeostasis of restored vegetation in the Loess Hilly Region[J]. Scientia Silvae Sinicae，2025（6）：61-74 张城伟，王兴，安可，等. 黄土丘陵区退耕植被叶片-土壤生态化学计量特征与植物内稳态差异[J]. 林业科学，2025（6）：61-74
[48] ELSER J J，FAGAN W F，KERKHOFF A J，et al. Biological stoichiometry of plant production： metabolism，scaling and ecological response to global change[J]. New Phytologist，2010，186（3）：593-608
[49] WANG Z F，ZHENG F L. Ecological stoichiometry of plant leaves，litter and soils in a secondary forest on China’s Loess Plateau[J]. PeerJ，2020，8：e10084
[50] LIU J H，ZHANG T. Response of plant characteristics and soil nutrients on grazing disturbance in typical grassland in Xilinguole[J]. Ecology and Environmental Sciences，2017，26（12）：2016-2023 刘佳慧，张韬. 放牧扰动对锡林郭勒典型草原植被特征及土壤养分的影响[J]. 生态环境学报，2017，26（12）：2016-2023
[51] AN Y，AN H，LI S B. Effects of grazing on ecological stoichiometric characteristics of soil and dominant plants in a desert steppe[J]. Acta Prataculturae Sinica，2018（12）：94-102 安钰，安慧，李生兵. 放牧对荒漠草原土壤和优势植物生态化学计量特征的影响[J]. 草业学报，2018（12）：94-102
[52] BAI L，CUI Y Y，LIU Z T，et al. Effects of grazing on leaf nutrient content and stoichiometric characteristics of dominant plants in a desert steppe[J]. Animal Husbandry and Feed Science，2021（6）：56-62 白柳，崔媛媛，刘倬彤，等. 放牧对荒漠草原优势植物叶片养分含量和化学计量特征的影响[J]. 畜牧与饲料科学，2021（6）：56-62

The Influence of Cervus elaphus kansuensis Grazing on Leaf-Soil Ecological Stoichiometry Characteristics of Plant Community in A Long-Term Enclosed Grassland of Ningxia

马鹿放牧对云雾山封育草地植物-土壤系统化学计量特征的影响

PDF

Knowledge

Abstract

Cite this article

share this article

References

Related Articles 15

Recommended Articles

Metrics

Comments

[1]	LIU Meng-di, ZHANG Zhen-jie, WANG Hong-mei, ZHAO Ya-nan, LI Ao-tian. Effects of Shrub Introduction on Soil C-N-P Stoichiometry and Their Trade-off with Water in a Desert Steppe [J]. Acta Agrestia Sinica, 2026, 34(3): 905-916.
[2]	WANG Xiao-li, WANG Qian, XU Hang, HE Ke-yan, WANG Jia-hao, SONG Mei-qi, JIN Ling, WEI Xiao-xing. Effects of Phosphorus Addition on Stoichiometric Traits and Nutrient Resorption Efficiency in Bluegrass Leaves [J]. Acta Agrestia Sinica, 2026, 34(3): 949-959.
[3]	FAN Xu-bin, YAN Bao-long, ZHANG Shu-han, GAO Ming-yue. Effects of Nitrogen Addition on Interspecific Association and Niche of Plants in Typical Steppe of Horqin [J]. Acta Agrestia Sinica, 2026, 34(3): 973-983.
[4]	MI Yu-qing, XU Hong-bin, ZHANG Lei, TANG Yue-kun, AO Hu-richa, WEN Du-rigen, LI Ai-lan. The Impact of Enclosure on Vegetation Ecological Niche and Interspecific Associations in Inner Mongolian Desert Steppe [J]. Acta Agrestia Sinica, 2026, 34(3): 1007-1016.
[5]	BI Xin, YU Tian-qi, SHI Meng-yuan, XIN Xiao-ping, ZHANG Zhi-tao, YAN Rui-rui, HAN Guo-dong. The Influences of Grazing Intensity on the Composition of Plant Communities and Ecological Niches in the Leymus chinensis Meadow Steppe [J]. Acta Agrestia Sinica, 2026, 34(2): 403-414.
[6]	LIU Zhao-qi, WANG Qi, ZHENG Jia-hua, ZHANG Feng, LI Shao-yu, YANG Li-shan, ZHANG Bin, QIAO Ji-rong, ZHAO Meng-li, XU Long-chao, WANG Ning-bin, ZHOU Qing-ge, DENG Yang-zhen, WANG Xi-yuan, CHEN Xin-li, JI Xiang, WANG Zhong-wu. Effects of Grazing Intensities on AMF Colonization of Stipa breviflora and their influencing factors in Desert Steppe [J]. Acta Agrestia Sinica, 2026, 34(2): 469-477.
[7]	CHEN Wen-mei, GAO Hui, HU Hai-na, HUANG Zhen-zhen, SUN Zhi, LI Hai-gang. The Effects of Soil Microorganisms on the Growth of Annual and Perennial Forages in the Desert Steppe [J]. Acta Agrestia Sinica, 2026, 34(1): 61-74.
[8]	DUAN Jun-guang, FANG Kai, CHU Jian-min, WANG Ying-xin, ZHANG Qi. Effects of Degradation Stages on the Relationship Between Species Diversity and Soil Nutrients in Typical Steppe [J]. Acta Agrestia Sinica, 2025, 33(9): 2890-2899.
[9]	YU Ya-nan, GE Nan, WANG Bo, LI Yu-wei, LYU Nan, WANG Chuan-qi, LI Xin. Ecological Stoichiometric Changes and Responses to Environment of Plant Root-Soil System in Desert Grassland Region of Central and Western Inner Mongolia [J]. Acta Agrestia Sinica, 2025, 33(9): 2921-2930.
[10]	JU Xin, WANG Xin-ya, WANG Bing-ying, HAN Guo-dong, WU Qian. Effects of Grazing Intensity on Plant Community Composition and Nutrient Quality of Forage in a Desert Steppe [J]. Acta Agrestia Sinica, 2025, 33(9): 2962-2972.
[11]	ZHANG Lei, ZHANG Yu-juan, SUN Shi-xian, XU Xue-bao, XING Yue, LIU Xiao-li, GAO Cui-ping, WANG Cheng-jie. Effects of Different Grazing Intensities on Soil Organic Carbon Content in Desert Steppe [J]. Acta Agrestia Sinica, 2025, 33(8): 2541-2547.
[12]	CHEN Meng-jiao, NIE Ze-xu, HE Feng-yuan, LIU Xin-yan, REN Fei. Spatial Variability Characteritics and Driving Mechanism of Vegetation Productivity of the Northern Grassland in Inner Mongolia， China [J]. Acta Agrestia Sinica, 2025, 33(8): 2677-2684.
[13]	ZHANG Li, YANG Xin-guo, WANG Lei, ZHANG Xue, QU Wen-jie, LIU Rong-guo, ZHANG Bo. Spatial Associations Analysis of Species in Sand-Fixing Community of Caragana korshinskii Based on Point Pattern [J]. Acta Agrestia Sinica, 2025, 33(6): 1886-1893.
[14]	JU Xin, WANG Wei, SHEN Ting-ting, WANG Yu-xin, HAN Guo-dong, WU Qian. Changes in Sheep Grazing Behavior under Different Grazing Intensities in Desert Steppe and Their Responses to Environmental Factors [J]. Acta Agrestia Sinica, 2025, 33(6): 1924-1933.
[15]	WANG Jia-jing, HE Si-teng, LIANG Xin-ran, YANG Meng-jiao, ZHANG Ya, YANG Zhi-xin, HE Yong-mei. The Impact of Euphorbia jolkinii Boiss. Invasion on the Soil-Plant Carbon， Nitrogen， and Phosphorus Ecological Stoichiometry in Subalpine Meadows of Northwest Yunnan [J]. Acta Agrestia Sinica, 2025, 33(5): 1523-1533.