[1] 李建东,方精云. 中国草地生态保障与食物安全战略研究(第三卷)[M]. 北京:科学出版社,2017:1-62 [2] 刘兴元,龙瑞军,尚占环. 青藏高原高寒草地生态系统服务功能的互作机制[J]. 生态学报,2012,32(24):7688-7697 [3] 白永飞,赵玉金,王扬,等. 中国北方草地生态系统服务评估和功能区划助力生态安全屏障建设[J]. 中国科学院院刊,2020,35(6):675-689 [4] 李通,崔丽珍,朱佳佩,等. 草地生态系统多功能性与可持续发展目标的实现[J]. 自然杂志,2021,43(2):149-156 [5] ASNER G P,ELMORE A J,OLANDER L P,et al. Grazing systems,ecosystem responses,and global change[J]. Annual Review of Environment and Resources,2004,29(1):261-299 [6] 张扬建,朱军涛,沈若楠,等. 放牧对草地生态系统影响的研究进展[J]. 植物生态学报,2020,44(5):553-564 [7] 周宸宇,杨晓渊,邵新庆,等. 不同退化程度高寒草甸植物物种多样性与生态系统多功能性关系[J]. 草地学报,2022,30(12):3410-3422 [8] LIU N,KAN H M,YANG G M,et al. Changes in plant,soil,and microbes in a typical steppe from simulated grazing:explaining potential change in soil C[J]. Ecological Monographs,2015,85(2):269-286 [9] MCNAUGHTON S J,BANYIKWA F F,MCNAUGHTON M M. Promotion of the cycling of diet-enhancing nutrients by African grazers[J]. Science,1998,278(5344):1798-1800 [10] YANG Y F,WU L W,LIN Q Y,et al. Responses of the functional structure of soil microbial community to livestock grazing in the Tibetan alpine grassland[J]. Global Change Biology,2013,19(2):637-648 [11] MUELLER P,GRANSE D,NOLTE S,et al. Top-down control of carbon sequestration:grazing affects microbial structure and function in salt marsh soils[J]. Ecological Applications,2017,27(5):1435-1450 [12] BARDGETT R D,WARDLE D A,YEATES G W. Linking above-ground and below-ground interactions:How plant responses to foliar herbivory influence soil organisms[J]. Soil Biology and Biochemistry,1998,30(14):1867-1878 [13] HAMILTON E W,FRANK D A. Can plants stimulate soil microbes and their own nutrient supply? Evidence from a grazing tolerant grass[J]. Ecology,2001,82(9):2397-2402 [14] 贾涛涛,廖李容,王杰,等. 基于Meta分析的放牧对黄土高原草地生态系统的影响[J]. 草地学报,2022,30(10):2772-2781 [15] KAUFFMAN J B,THORPE A S,BROOKSHIRE E. Livestock exclusion and belowground ecosystem responses in riparian meadows of eastern Oregon[J]. Ecological Applications,2004,14(6):1671-1679 [16] KOHLER F,HAMELIN J,GILLET F G,et al. Soil microbial community changes in wooded mountain pastures due to simulated effects of cattle grazing[J]. Plant and Soil,2005,278(1):327-340 [17] WANG D L,WANG L,LIU J S,et al. Grassland ecology in China:perspectives and challenges[J]. Frontiers of Agricultural Science and Engineering,2018,5(1):24-43 [18] MCINTIRE C D,COLBY J A. A Hierarchical Model of Lotic Ecosystems[J]. Ecological Monographs,1978,48(2):167-190 [19] MCNAUGHTON S J. Ecology of a grazing ecosystem:The Serengeti[J]. Ecological Monographs,1985,55(3):259-294 [20] AUGUSTINE D J,FRELICH L E,JORDAN P A. Evidence for two alternate stable states in an ungulate grazing system[J]. Ecological Applications,1998,8(4):1260-1269 [21] MILCHUNAS D G,LAUENROTH W K. Quantitative effects of grazing on vegetation and soils over a global range of environments[J]. Ecological Monographs,1993,63(4):327-366 [22] NIKLAUS P A,ALPHEI J,KAMPICHLER C,et al. Interactive effects of plant species diversity and elevated CO2 on soil biota and nutrient cycling[J]. Ecology,2007,88(12):3153-3163 [23] BAKONYI G,NAGY P. Temperature-and moisture-induced changes in the structure of the nematode fauna of a semiarid grassland-patterns and mechanisms[J]. Global Change Biology,2010,6(6):697-707 [24] COLE L,STANDDON P L,SLEEP D,et al. Soil animals influence microbial abundance,but not plant-microbial competition for soil organic nitrogen[J]. Functional Ecology,2004,18(5):631-640 [25] AKINOLA M O,BUCKLAND T S M. Soil seed bank of an upland calcareous grassland after 6 years of climate and management manipulations[J]. Journal of Applied Ecology,1998,35(4):544-552 [26] LI J J,HUANG Y,XU F W,et al. Responses of growing-season soil respiration to water and nitrogen addition as affected by grazing intensity[J]. Functional Ecology,2018,32(7):1890-1901 [27] WANG C L,ZHANG R,VILONEN L,et al. Grazing and nitrogen addition restructure the spatial heterogeneity of soil microbial community structure and enzymatic activities[J]. Functional Ecology,2021,35(12):2763-2777 [28] 王岭,张敏娜,徐曼,等. 草地多功能提升的多样化家畜放牧理论及应用[J]. 科学通报,2021,66(30):3791-3798 [29] 杨殿林,韩国栋,胡跃高,等. 放牧对贝加尔针茅草原群落植物多样性和生产力的影响[J]. 生态学杂志,2006(12):1470-1475 [30] YAN R R,XIN X P,YAN Y C,et al. Impacts of differing grazing rates on canopy structure and species composition in Hulunber meadow steppe[J]. Rangeland Ecology and Management,2015,68(1):54-64 [31] LI W,WU G L,ZHANG G F,et al. The maintenance of offspring diversity in response to land use:sexual and asexual recruitment in an alpine meadow on the Tibetan Plateau[J]. Nordic Journal of Botany,2011,29(1):81-86 [32] PENG J T,LIANG C Z,NIU Y M,et al. Moderate grazing promotes genetic diversity of Stipa species in the Inner Mongolian steppe[J]. Landscape Ecology,2015,30(9):1783-1794 [33] GAO J J,CARMEL Y. Can the intermediate disturbance hypothesis explain grazing-diversity relations at a global scale?[J]. Oikos,2020,129(4):493-502 [34] 刘玉祯,孙彩彩,刘文亭,等. 高寒草地植物群落关键种对不同放牧家畜组合放牧的响应[J]. 生态学报,2022,42(18):7529-7540 [35] WILSON P J,THOMPSON K,HODGSON J G. Specific leaf area and leaf dry matter content as alternative predictors of plant strategies[J]. New Phytologist,2010,143(1):155-162 [36] 王晓芳,马红彬,刘杰,等. 放牧对草原植物功能性状影响研究进展[J]. 应用生态学报,2022,33(2):569-576 [37] WANG D L,DU J,ZHANG B T,et al. Grazing intensity and phenotypic plasticity in the clonal grass Leymus chinensis[J]. Rangeland Ecology and Management,2017,70(6):740-747 [38] BAI W M,FANG Y,ZHOU M,et al. Heavily intensified grazing reduces root production in an Inner Mongolia temperate steppe[J]. Agriculture,Ecosystems and Environment,2015,1(200):143-150 [39] 崔猛,冯媛媛,王新宇,等. 不同放牧方式对松嫩草地地下、地上生物量及其分配比例的影响[J]. 东北师大学报(自然科学版),2021,53(4):144-150 [40] 许宏斌,辛晓平,宝音陶格涛,等. 放牧对呼伦贝尔羊草草甸草原生物量分布的影响[J]. 草地学报,2020,28(3):768-774 [41] MA L,GUO C,LU X,et al. Soil moisture and land use are major determinants of soil microbial community composition and biomass at a regional scale in northeastern China[J]. Biogeosciences,2015,12(8):2585-2596 [42] 吴东辉,尹文英,卜照义. 松嫩草原中度退化草地不同植被恢复方式下土壤线虫的群落特征[J]. 生态学报,2008,28(1):1-12 [43] 王永宏,田黎明,艾鷖,等. 短期牦牛放牧对青藏高原高寒草地土壤真菌群落的影响[J]. 草业学报,2022,31(10):41-52 [44] 郑佳华,赵萌莉,王琪,等. 放牧和刈割对大针茅草原土壤微生物群落结构及多样性的影响[J]. 生态学报,2022,42(12):4998-5008 [45] 赵熠. 不同放牧强度对短花针茅荒漠草原土壤线虫群落组成及多样性的影响[D]. 呼和浩特:内蒙古农业大学,2021:1-42 [46] 王亚东. 放牧家畜类型和强度对内蒙古典型草原地表节肢动物群落的影响[D]. 呼和浩特:内蒙古大学,2021:1-36 [47] QI Q,ZHAO M X,WANG S P,et al. The biogeographic pattern of microbial functional genes along an altitudinal gradient of the Tibetan Pasture[J]. Frontiers in Microbiology,2017,13(8):976 [48] 胡靖,钱秀娟,刘长仲. 放牧模式对高山草地蝗虫群落生物多样性的影响及其作用机制[J]. 植物保护学报,2021,48(1):212-220 [49] 潘多锋. 青藏高原高寒草甸草食动物间的相互作用关系及机制[D]. 长春:东北师范大学,2019:26-53 [50] ARIANNE J C,JAMES J E,COLLEEN F F,et al. Heavy livestock grazing promotes locust outbreaks by lowering plant nitrogen content[J]. Science,2012,335(6067):467-469 [51] ZHONG Z W,WANG D L,ZHU H,et al. Positive interactions between large herbivores and grasshoppers,and their consequences for grassland plant diversity[J]. Ecology,2016,95(4):1055-1064 [52] ZHU H,WANG D L,WANG L,et al. The effects of large herbivore grazing on meadow steppe plant and insect diversity[J]. Journal of Applied Ecology,2012,49(5):1075-1083 [53] ZHU H,QU Y K,ZHANG D,et al. Impacts of grazing intensity and increased precipitation on a grasshopper assemblage (Orthoptera:Acrididae) in a meadow steppe[J]. Ecological Entomology,2017,42(4):458-468 [54] LI W H,ZHAN S X,LAN Z C,et al. Scale-dependent patterns and mechanisms of grazing-induced biodiversity loss:evidence from a field manipulation experiment in semiarid steppe[J]. Landscape Ecology,2015,30(9):1751-1765 [55] 姚国征,高永,杨婷婷,等. 放牧对小针茅荒漠草原枯落物及植被生产力的影响[J]. 干旱区资源与环境,2016,30(10):93-97 [56] YAN L,ZHOU G S,ZHANG F. Effects of different grazing intensities on grassland production in China:A meta-analysis[J]. PloS One,2013,8(12):e81446 [57] 李勤奋,韩国栋,卫智军,等. 放牧制度对短花针茅草原植物群落的影响[J]. 农业现代化研究,2002,89(3):192-196 [58] SCHIMEL J P,BENNETT J. Nitrogen mineralization:challenges of a changing paradigm[J]. Ecology,2004,85(3):591-602 [59] BAI Y F,WU J G,CLARK M C,et al. Grazing alters ecosystem functioning and C:N:P stoichiometry of grasslands along a regional precipitation gradient[J]. Journal of Applied Ecology,2012,49(6):1204-1215 [60] SHAN Y M,CHEN D M,GUAN X X,et al. Seasonally dependent impacts of grazing on soil nitrogen mineralization and linkages to ecosystem functioning in Inner Mongolia grassland[J]. Soil Biology and Biochemistry,2011,43(9):1943-1954 [61] LIU C,WANG L,SONG X X,et al. Towards a mechanistic understanding of the effect that different species of large grazers have on grassland soil N availability[J]. Journal of Ecology,2017,106(1):357-366 [62] 刘玉祯,刘文亭,杨晓霞,等. 放牧对全球草地生态系统碳氮磷化学计量特征影响的Meta分析[J]. 应用生态学报,2022,33(5):1251-1259 [63] WANG S P,ANDREAS W,ZHANG Z C,et al. Management and land use change effects on soil carbon in northern China's grasslands:a synthesis[J]. Agriculture Ecosystems and Environment,2011,142(3-4):329-340 [64] LI C L,HAO X Y,ZHAO M L,et al. Influence of historic sheep grazing on vegetation and soil properties of a desert steppe in Inner Mongolia[J]. Agriculture Ecosystems & Environment,2008,128(2):109-116 [65] WANG J L,LIU Y Z,CAO W X,et al. Effects of grazing exclusion on soil respiration components in an alpine meadow on the north-eastern Qinghai-Tibet Plateau[J]. Catena,2020,194(7):104750 [66] WANG L,WANG D L,HE Z B,et al. Mechanisms linking plant species richness to foraging of a large herbivore[J]. Journal of Applied Ecology,2010,47(4):868-875 [67] WANG L,WANG D L,BAI Y G,et al. Spatially complex neighboring relationships among grassland plant species as an effective mechanism of defense against herbivory[J]. Oecologia,2010,164(1):193-200 [68] NIU S L,LI Z X,XIA J Y,et al. Climatic warming changes plant photosynthesis and its temperature dependence in a temperate steppe of northern China[J]. Environmental and Experimental Botany,2008,63(1-3):91-101 [69] CHI Y G,XU M,SHEN R C,et al. Acclimation of foliar respiration and photosynthesis in response to experimental warming in a temperate steppe in Northern China[J]. PloS One,2013,8(2):e56482 [70] YANG H J,WU M G,LIU W X,et al. Community structure and composition in response to climate change in a temperate steppe[J]. Global Change Biology,2015,17(1):452-465 [71] YANG H J,LI Y,WU M Y,et al. Plant community responses to nitrogen addition and increased precipitation:the importance of water availability and species traits[J]. Global Change Biology,2011,17(9):2936-2944 [72] BAI W M,WANG Z W,CHEN Q S,et al. Spatial and temporal effects of nitrogen addition on root life span of Leymus chinensis in a typical steppe of Inner Mongolia[J]. Functional Ecology,2008,22(4):583-591 [73] LIN D L,XIA J Y,WAN S Q. Climate warming and biomass accumulation of terrestrial plants:A meta-analysis[J]. New Phytologist,2010,188(1):187-198 [74] XU Z W,REN H Y,LI M H,et al. Environmental changes drive the temporal stability of semi-arid natural grasslands through altering species asynchrony[J]. Journal of Ecology,2015,103(5):1308-1316 [75] ZHANG N L,WAN S Q,GUO J X,et al. Precipitation modifies the effects of warming and nitrogen addition on soil microbial communities in northern Chinese grasslands[J]. Soil Biology and Biochemistry,2015,89(10):12-23 [76] WU T J,SU F L,HAN H Y,et al. Responses of soil microarthropods to warming and increased precipitation in a semiarid temperate steppe[J]. Applied Soil Ecology,2014,84(12):200-207 [77] SONG M,JING S S,ZHOU Y Q,et al. Dynamics of soil nematode communities in wheat fields under different nitrogen management in Northern China Plain[J]. European Journal of Soil Biology,2015,71(11-12):13-20 [78] ZHU H,ZOU X H,WANG D L,et al. Responses of community-level plant-insect interactions to climate warming in a meadow steppe[J]. Scientific Reports,2015,5:18654 [79] LIU H Y,Wang H,Li N,et al. Phenological mismatches between above- and belowground plant responses to climate warming[J]. Nature Climate Change,2022,12:97-102 [80] LIANG M W,LIANG C,HAUTIER Y,et al. Grazing-induced biodiversity loss impairs grassland ecosystem stability at multiple scales[J]. Ecology Letters,2021,24(10):2054-2064 [81] REN H,EVINER V T,GUI W,et al. Livestock grazing regulates ecosystem multifunctionality in semi-arid grassland[J]. Functional Ecology,2018,32(12):2790-2800 [82] 徐万玲. 氮沉降、放牧和极端降水对羊草草地N2O排放的影响机制研究[D]. 长春:东北师范大学,2021:21-45 [83] WANG L,DELGADO-BAQUERIZO M,WANG D L,et al. Diversifying livestock promotes multidiversity and multifunctionality in managed grasslands[J]. Proceedings of the National Academy of Sciences,2019,116(13):6187-6192 |