中国森林大型真菌重要类群多样性和系统学研究
Diversity and systematics of the important macrofungi in Chinese forests
第一联系人: △ Contributed equally as the first authors.
责任编辑: 韩丽
收稿日期: 2021-01-22 接受日期: 2021-02-2 网络出版日期: 2021-04-22
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Corresponding authors:
Received: 2021-01-22 Accepted: 2021-02-2 Online: 2021-04-22
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作者简介 About authors
戴玉成,北京林业大学教授,博士生导师,国家杰出青年基金获得者主要从事菌物分类与分子系统学、多样性与生态学、森林病理学、菌物资源与利用等领域的研究,发现和发表真菌新科3个,新属40个,新种400余个,森林新病原真菌72个,发表论文430余篇,出版专著10部,获发明专利12项,获得省部级科技奖励9项现任中国菌物学会副理事长,《菌物学报》主编,《菌物研究》《北京林业大学学报》和《生物技术通报》副主编 , E-mail:yuchengd@yahoo.com
杨祝良,中国科学院昆明植物研究所研究员,博士生导师,国家杰出青年基金获得者主要从事真菌系统学研究,现任中国科学院东亚植物多样性与生物地理学重点实验室主任,兼任《FungalDiversity》主编、《菌物学报》和《菌物研究》副主编,兼任国际真菌命名法委员会委员,发表学术论文230余篇,出版专著14部,发表24个新属和292个新种曾获省部级科研成果一、二等奖数次,获全国先进工作者称号、全国五一劳动奖章、中国菌物学会戴芳澜杰出成就奖及美国真菌学会荣誉会员奖等 , E-mail:fungi@mail.kib.ac.cn
大型真菌主要为担子菌门的真菌和少数为子囊菌门的真菌,该类真菌具有重要的经济价值和生态功能,主要生长在森林生态系统中。30年来作者对我国几乎所有类型森林生态中的大型真菌进行了系统调查和采集,共采集标本11.2万号。基于对这些材料的形态学及分子系统学研究,并结合生态学和生物地理学特征,共鉴定出中国森林大型真菌4 250种,隶属于担子菌门和子囊菌门的21个目,发现和发表2个新科、4个新亚科、69个新属和885个新种。云南省是我国森林大型真菌最丰富的省份,描述于该省的新种有314种,占作者发表的全部中国新种的35%。这些研究为深入认识全球大型真菌物种多样性提供了中国的贡献,更新了我国重要食药用菌名称,揭示了我国毒蘑菇多样性基本特征,系统论述了我国森林病原菌的物种多样性,为资源利用、森林健康和保护提供了科学依据;论述了森林大型真菌代表性类群在种和属级水平的起源和演化,为今后开展重要类群科级、目级甚至纲级的系统进化关系提供了重要数据。
关键词:
Macrofungi, as an important component in forest ecosystems, consist of most members of Basidiomycota and some members of Ascomycota, having important economical value and ecological functions. Extensive field investigations have been carried out in almost whole types of the Chinese forests during the past 30 years, and 112 000 specimens were collected. Based on morphological examination and phylogenetic analyses in combination with ecology and biogeography, 4 250 species belonging to 21 orders in Baidiomycota and Ascomycota were identified, including two new families, four new subfamilies, 69 new genera and 885 new species. Yunnan Province is the richest in macrofungal diversity among provinces or regions in China, and 314 new species were described from this province, accounting for 35% of all the new species described from China by the authors. Our studies have made contributions to deepening the understanding of global diversity of macrofungi. The names of some important Chinese medicinal fungi were revised, the diversity characteristics of Chinese poisonous mushrooms were revealed, and the pathogenetic wood-decaying species were ascertained. These data improved our knowledge on utilization of natural resources and protection of forest health. Based on molecular evidences, the origin of some forest representative fungal genera or species complex were deduced, and their dispersal and speciation were discussed, for the purposes of providing some data for evolutionary study at level of family, order or class of macrofungi henceforth.
Keywords:
本文引用格式
戴玉成, 杨祝良, 崔宝凯, 吴刚, 袁海生, 周丽伟, 何双辉, 葛再伟, 吴芳, 魏玉莲, 员瑗, 司静.
DAI Yu-Cheng, YANG Zhu-Liang, CUI Bao-Kai, WU Gang, YUAN Hai-Sheng, ZHOU Li-Wei, HE Shuang-Hui, GE Zai-Wei, WU Fang, WEI Yu-Lian, YUAN Yuan, SI Jing.
大型真菌通常是指肉眼可见、徒手可摘的真菌,主要为担子菌门Basidiomycota、少数为子囊菌门Ascomycota真菌。该类真菌具有重要的经济价值:目前绝大多数野生或栽培的食用菌,包括松茸Tricholoma matsutake (S. Ito & S. Imai) Singer、香菇Lentinula edodes (Berk.) Pegler和黑木耳Auricularia heimuer F. Wu et al.等都是大型真菌(戴玉成等 2010);同样我国绝大多数药用真菌,包括灵芝Ganoderma lingzhi Sheng H. Wu et al.和猴头Hericium erinaceus (Bull.) Pers.等也都属于大型真菌(戴玉成和杨祝良 2008)。该类真菌还具有重要的生态价值:最主要的外生菌根真菌如牛肝菌科Boletaceae和红菇科Russulaceae,最重要的木材降解菌如多孔菌科Polyporaceae和锈革孔菌科Hymenochaetaceae等科的种类也都是大型真菌(李玉等 2015)。该类真菌也影响森林健康:一些重要的森林病原真菌,包括异担子菌属Heterobasidion Bref.和蜜环菌属Armillaria (Fr.) Staude等的种类能够造成重大森林病害(戴玉成 2012)。该类真菌还影响人类健康:一些种类是剧毒蘑菇,据国家食品安全风险评估中心统计,2020年我国有约8 000人误食毒蘑菇,74人死亡,准确鉴定这些剧毒蘑菇对预防和治疗蘑菇中毒具有重大意义。因此,大型真菌在生态系统中对促进植物生长、维护生态系统的稳定性及物质循环和能量流动发挥不可替代的作用,也在一定程度上影响森林和人类的健康。
在用双名法最早描述的植物中,就包括了木蹄层孔菌Fomes fomentarius (L.) Fr. ≡ Boletus fomentarius L.、火木层孔菌Phellinus igniarius (L.) Quél. ≡ Boletus igniarius L.和点柄乳牛肝菌Suillus granulatus (L.) Roussel ≡ Boletus granulatus L.等大型真菌(Linnaeus 1753)。虽然我国从约2 000年前就开始对大型真菌进行了诸多的记载和药用功能描述,但以双名法规对我国大型真菌进行分类学研究始于18世纪。一些外国传教士在中国进行植物标本采集时也一并采集了大型真菌,并进行了系列报道。最早以我国材料记载的大型真菌为1782年发表的五棱散尾鬼笔Lysurus mokusin (L.) Fr. ≡ Phallus mokusin L.,其标本是由法国传教士西博特(P.M. Cibot)在中国采集的(戴芳澜 1979a)。我国著名真菌学家邓叔群院士是中国大型真菌研究的先驱,他在20世纪30年代对我国大型真菌进行了系列报道(Teng 1934,1935,1936,1939)。1979年出版的《中国真菌总汇》中记载了大型真菌种级分类单元名称2 020个,其中担子菌1 820种(戴芳澜 1979b)。
尽管我国科学家对中国大型真菌资源进行了系列研究,但若与植物多样性研究相比,大型真菌多样性研究仍然严重滞后,物种多样性认识严重不足,诸多系统学问题有待解决。特别是由于大型真菌化石较少,多数类群的起源和演化问题处于探索阶段。在各种生态系统中,森林生态系统植被区系组成复杂、生境多样,孕育了丰富的大型真菌资源。因此森林生态系统的大型真菌物种多样性最高,但过去的研究还不够充分和深入。北京林业大学、中国科学院昆明植物研究所和中国科学院沈阳应用生态研究所的学者30年来对全国所有省区市550多个国家公园、自然保护区、森林公园和林场的森林大型真菌多样性和资源进行了广泛考察,获得标本近11.2万号,保藏于上述3个单位的标本馆中(BJFC、KUN和IFP)。基于对上述材料的系统学研究,我们取得的主要研究结果总结如下。
1 发现中国森林大型真菌4 250种
基于标本的形态学、生态学或分子系统学研究数据,共鉴定出4 250种中国森林大型真菌,隶属于担子菌门的伞菌目Agaricales、淀粉孢革菌目Amylocorticiales、阿太菌目Atheliales、木耳目Auriculariales、牛肝菌目Boletales、鸡油菌目Cantharellales、伏革菌目Corticiales、花耳目Dacrymycetales、地星目Geastrales、褐褶菌目Gloeophyllales、钉菇目Gomphales、锈革孔菌目Hymenochaetales、鬼笔目Phallales、多孔菌目Polyporales、红菇目Russulales、蜡壳菌目Sebacinales、革菌目Thelephorales、糙孢菌目Trechisporales和银耳目Tremellales,以及子囊菌门的肉座菌目Hypocreales和盘菌目Pezizales等21个目中(Yang 1997;杨祝良 2005,2015;Dai 2011,2012;Wu et al. 2014,2016;Cai et al. 2016;杨祝良等 2017,2019;Cui et al. 2018,2019;戴玉成 2018;Han et al. 2019;Li et al. 2020)。
2 发现森林大型真菌重要类群新分类单元960个
利用形态特征、分子数据和生态习性等多相证据,发现和发表2个新科、4个新亚科、69个新属和885个新种,具体情况如下。
新科
1.Fragiliporiaceae Y.C. Dai, B.K. Cui & C.L. Zhao, Fungal Diversity 70: 119, 2015
2.Neoantrodiellaceae Y.C. Dai, B.K. Cui, Jia J. Chen & H.S. Yuan, Fungal Diversity 75: 228, 2015
新亚科
1.Austroboletoideae G. Wu & Zhu L. Yang, Fungal Diversity 69: 104, 2014
2.Chalciporoideae G. Wu & Zhu L. Yang, Fungal Diversity 69: 105, 2014
3.Leccinoideae G. Wu & Zhu L. Yang, Fungal Diversity 69: 105, 2014
4.Zangioideae G. Wu, Y.C. Li & Zhu L. Yang, Fungal Diversity 69: 106, 2014
新属
1.Amaropostia B.K. Cui, L.L. Shen & Y.C. Dai, Persoonia 42: 110, 2019
2.Amylosporia B.K. Cui, C.L. Zhao & Y.C. Dai, Fungal Diversity 97: 164, 2019
3.Baorangia G. Wu & Zhu L. Yang, Fungal Diversity 81: 2, 2016
4.Calcipostia B.K. Cui, L.L. Shen & Y.C. Dai, Persoonia 42: 111, 2019
5.Cerarioporia F. Wu, L.W. Zhou & J. Si, Phytotaxa 280: 57, 2016
6.Chiua Yan C. Li & Zhu L. Yang, Fungal Diversity 81: 76, 2016
7.Cibaomyces Zhu L. Yang et al., Phytotaxa 160: 203, 2014
8.Coniferiporia L.W. Zhou & Y.C. Dai, Fungal Biology 120: 993, 2016
9.Crassisporus B.K. Cui & Xing Ji, MycoKeys 57: 69, 2019
10.Cylindrosporus L.W. Zhou & Y.C. Dai, Phytotaxa 219: 279, 2015
11.Cystidiopostia B.K. Cui, L.L. Shen & Y.C. Dai, Persoonia 42: 118, 2019
12.Dacryoscyphus R. Kirschner & Zhu L. Yang, Antonie van Leeuwenhoek 87: 331, 2005
13.Datroniella B.K. Cui, Hai J. Li & Y.C. Dai, Persoonia 32: 172, 2014
14.Dentipellicula Y.C. Dai & L.W. Zhou, Mycologia 105: 639, 2013
15.Dentipellopsis Y.C. Dai & L.W. Zhou, Mycologia 105: 642, 2013
16.Dextrinoporus H.S. Yuan, Mycological Progress 17: 774, 2018
17.Donkioporiella L.W. Zhou, Cryptogamie Mycologie 37: 439, 2016
18.Echinodontiellum S.H. He & Nakasone, Mycologia 109: 571, 2017
19.Flammeopellis Y.C. Dai, B.K. Cui & C.L. Zhao, Mycological Progress 13: 776, 2014
20.Fragifomes B.K. Cui, M.L. Han & Y.C. Dai, Fungal Diversity 80: 360, 2016
21.Fragiliporia Y.C. Dai, B.K. Cui & C.L. Zhao, Fungal Diversity 70: 121, 2015
22.Fulvoderma L.W. Zhou & Y.C. Dai, Mycologia 110: 876, 2018
23.Fuscopostia B.K. Cui, L.L. Shen & Y.C. Dai, Persoonia 42: 118, 2019
24.Geliporus Yuan Yuan, Jia J. Chen & S.H. He, Mycoscience 58: 200, 2017
25.Grammatus H.S. Yuan & C. Decock, MycoKeys 35: 32, 2018
26.Hispidaedalea Y.C. Dai & S.H. He, Mycological Progress 13: 835, 2014
27.Hourangia Xue T. Zhu & Zhu L. Yang, Mycological Progress 14: 37/36, 2015
28.Hymenoboletus Yan C. Li & Zhu L. Yang, Fungal Diversity 81: 100, 2016
29.Hymenochaetopsis S.H. He & Jiao Yang, Mycological Progress 15: 13, 2016
30.Lanmaoa G. Wu, Zhu L. Yang, Halling, Fungal Diversity 81: 7, 2016
31.Larssoniporia Y.C. Dai, Jia J. Chen & B.K. Cui, Persoonia 37: 26, 2016
32.Lauriliella S.H. He & Nakasone, Mycologia 109: 571, 2017
33.Leifiporia Y.C. Dai, F. Wu & C.L. Zhao, Mycological Progress 15: 802, 2016
34.Liangia H. Yu, Y.B. Wang, Y. Wang, Z.H. Chen & Zhu L. Yang, Fungal Diversity 103: 22, 2020
35.Limacellopsis Zhu L. Yang, Qing Cai & Yang-Yang Cui, Biosystematics and Ecology Series 34: 366, 2018
36.Luteoporia F. Wu, Jia J. Chen & S.H. He, Phytotaxa 263: 37, 2016
37.Megasporia B.K. Cui, Y.C. Dai & Hai J. Li, Mycologia 105: 369, 2013
38.Megasporoporiella B.K. Cui, Y.C. Dai & Hai J. Li, Mycologia 105: 377, 2013
39.Melanoderma B.K. Cui & Y.C. Dai, Mycotaxon 116: 297, 2011
40.Murinicarpus B.K. Cui & Y.C. Dai, Fungal Diversity 97: 255, 2019
41.Neoantrodiella Y.C. Dai, B.K. Cui, Jia J. Chen & H.S. Yuan, Fungal Diversity 75: 228, 2015
42.Neodatronia B.K. Cui, Hai J. Li & Y.C. Dai, Persoonia 32: 177, 2014
43.Neofomitella Y.C. Dai, Hai J. Li & Vlasák, Mycotaxon 129: 12, 2015
44.Neomensularia F. Wu, L.W. Zhou & Y.C. Dai, Mycologia 108: 892, 2016
45.Niveoporofomes B.K. Cui, M.L. Han & Y.C. Dai, Fungal Diversity 80: 360, 2016
46.Ovipoculum Zhu L. Yang & R. Kirschner, Fungal Diversity 43: 58, 2010
47.Parvixerocomus G. Wu & Zhu L. Yang, Fungal Diversity 81: 11, 2016
48.Phellinopsis Y.C. Dai, Fungal Diversity 45: 309, 2010
49.Phlebiporia Jia J. Chen, B.K. Cui & Y.C. Dai, Mycological Progress 13: 568, 2014
50.Piptoporellus B.K. Cui, M.L. Han & Y.C. Dai, Fungal Diversity 80: 361, 2016
51.Pseudofibroporia Yuan Y. Chen & B.K. Cui, Mycological Progress 16: 527, 2017
52.Pseudowrightoporia Y.C. Dai, Jia J. Chen & B.K. Cui, Persoonia 37: 28, 2016
53.Radulotubus Y.C. Dai, S.H. He & C.L. Zhao, Nova Hedwigia 103: 270, 2016
54.Rhodofomitopsis B.K. Cui, M.L. Han & Y.C. Dai, Fungal Diversity 80: 365, 2016
55.Rigidotubus J. Song, Y.C. Dai & B.K. Cui, Phytotaxa 333: 263, 2018
56.Rubellofomes B.K. Cui, M.L. Han & Y.C. Dai, Fungal Diversity 80: 366, 2016
57.Rubroboletus Kuan Zhao & Zhu L. Yang, Phytotaxa 188: 67, 2014
58.Rugiboletus G. Wu & Zhu L. Yang, Fungal Diversity 81: 12, 2016
59.Sanghuangporus Sheng H. Wu, L.W. Zhou & Y.C. Dai, Fungal Diversity 77: 340, 2016
60.Sanguinoderma Y.F. Sun, D.H. Costa & B.K. Cui, Persoonia 44: 224, 2020
61.Taiwanofungus Sheng H. Wu et al., Fungal Science, Taipei 19: 110, 2004
62.Tengioboletus G. Wu & Zhu L. Yang, Fungal Diversity 81: 146, 2016
63.Tricholyophyllum Qing Cai, G. Kost & Zhu L. Yang, Mycosystema 39: 1734, 2020
64.Tropicoporus L.W. Zhou, Y.C. Dai & Sheng H. Wu, Fungal Diversity 77: 341, 2016
65.Tylocinum Yan C. Li & Zhu L. Yang, Fungal Diversity 81: 147, 2016
66.Ungulidaedalea B.K. Cui, M.L. Han & Y.C. Dai, Fungal Diversity 80: 366, 2016
67.Wrightoporiopsis Y.C. Dai, Jia J. Chen & B.K. Cui, Persoonia 37: 31, 2016
68.Yuchengia B.K. Cui & K.T. Steffen, Nordic J Botany 31: 333, 2013
69.Zangia Yan C. Li et Zhu L. Yang, Fungal Diversity 49: 129, 2011
新种及模式产地
1.Abundisporus mollissimus B.K. Cui & C.L. Zhao, 海南
2.Abundisporus quercicola Y.C. Dai, 云南
3.Acanthobasidium bambusicola L.D. Dai & S.H. He, 湖南
4.Agrocybe salicaceicola Zhu L. Yang, M. Zang & X.X. Liu, 云南
5.Albatrellus piceiphilus B.K. Cui & Y.C. Dai, 甘肃
6.Albatrellus zhuangii Y.C. Dai & Juan Li, 湖北
7.Aleurodiscus bambusinus S.H. He & Y.C. Dai, 江西
8.Aleurodiscus effusus L.D. Dai & S.H. He, 湖南
9.Aleurodiscus isabellinus S.H. He & Y.C. Dai, 云南
10.Aleurodiscus ryvardenii S.H. He & Y.C. Dai, 云南
11.Aleurodiscus subroseus S.H. He & Y.C. Dai, 广西
12.Aleurodiscus tenuissimus L.D. Dai & S.H. He, 海南
13.Aleurodiscus tropicus L.D. Dai & S.H. He, 海南
14.Aleurodiscus verrucosporus S.H. He, 福建
15.Amanita albidostipes Yang-Yang Cui, Qing Cai & Zhu L. Yang, 云南
16.Amanita alboumbelliformis Yang-Yang Cui, Qing Cai & Zhu L. Yang, 云南
17.Amanita altipes Zhu L. Yang, M. Weiss & Oberw., 云南
18.Amanita aspericeps Yang-Yang Cui, Qing Cai & Zhu L. Yang, 广东
19.Amanita atrofusca Zhu L. Yang, 四川
20.Amanita avellaneifolia Zhu L. Yang, Yang-Yang Cui & Qing Cai, 云南
21.Amanita brunneofuliginea Zhu L. Yang, 云南
22.Amanita brunneolimbata Zhu L. Yang, Yang-Yang Cui & Qing Cai, 广东
23.Amanita brunneomaculata Yang-Yang Cui, Qing Cai & Zhu L. Yang, 云南
24.Amanita brunneostrobilipes Zhu L. Yang, Yang-Yang Cui & Qing Cai, 海南
25.Amanita caojizong Zhu L. Yang, Yang-Yang Cui and Qing Cai, 云南
26.Amanita changtuia Yang-Yang Cui, Qing Cai & Zhu L. Yang, 黑龙江
27.Amanita chiui Yang-Yang Cui, Qing Cai & Zhu L. Yang, 四川
28.Amanita cinereoradicata Yang-Yang Cui, Qing Cai & Zhu L. Yang, 云南
29.Amanita cingulata J.W. Liu & Zhu L. Yang, 云南
30.Amanita citrinoannulata Yang-Yang Cui, Qing Cai & Zhu L. Yang, 重庆
31.Amanita citrinoindusiata Zhu L. Yang, Yang-Yang Cui & Qing Cai, 云南
32.Amanita detersa Zhu L. Yang, Yang-Yang Cui & Qing Cai, 云南
33.Amanita elliptica Qing Cai, Yang-Yang Cui & Zhu L. Yang, 广东
34.Amanita exitialis Zhu L. Yang & T.H. Li, 广东
35.Amanita flavopantherina Yang-Yang Cui, Qing Cai & Zhu L. Yang, 云南
36.Amanita flavosquamosa Yang-Yang Cui, Qing Cai & Zhu L. Yang, 云南
37.Amanita franzii Zhu L. Yang, Yang-Yang Cui & Qing Cai, 云南
38.Amanita fuligineoides P. Zhang & Zhu L. Yang, 湖南
39.Amanita fuscoflava Zhu L. Yang, Yang-Yang Cui & Qing Cai, 海南
40.Amanita griseofolia Zhu L. Yang, 云南
41.Amanita griseopantherina Yang-Yang Cui, Qing Cai & Zhu L. Yang, 西藏
42.Amanita griseorosea Qing Cai, Zhu L. Yang & Yang-Yang Cui, 广东
43.Amanita griseoumbonata Yang-Yang Cui, Qing Cai & Zhu L. Yang, 黑龙江
44.Amanita griseoverrucosa Zhu L. Yang ex Zhu L. Yang, 云南
45.Amanita innatifibrilla Zhu L. Yang ex Zhu L. Yang, Yang-Yang Cui & Qing Cai, 海南
46.Amanita lanigera Yang-Yang Cui, Qing Cai & Zhu L. Yang, 云南
47.Amanita lignitincta Zhu L. Yang ex Yang-Yang Cui, Qing Cai & Zhu L. Yang, 云南
48.Amanita liquii Zhu L. Yang, M. Weiss & Oberw., 云南
49.Amanita luteofolia Yang-Yang Cui, Qing Cai & Zhu L. Yang, 湖北
50.Amanita macrocarpa W.Q. Deng, T.H. Li & Zhu L. Yang, 广东
51.Amanita melleialba Zhu L. Yang, Q. Cai & Yang Y. Cui, 云南
52.Amanita minutisquama Yang-Yang Cui, Qing Cai & Zhu L. Yang, 海南
53.Amanita molliuscula Qing Cai, Zhu L. Yang & Yang-Yang Cui, 陕西
54.Amanita neocinctipes Zhu L. Yang, Yang-Yang Cui & Qing Cai, 广东
55.Amanita ochracea (Zhu L. Yang) Yang-Yang Cui, Qing Cai & Zhu L. Yang, 云南
56.Amanita olivaceofusca Yang-Yang Cui, Qing Cai & Zhu L. Yang, 云南
57.Amanita orienticrocea Zhu L. Yang, Yang-Yang Cui & Qing Cai, 云南
58.Amanita orientifulva Zhu L. Yang, M. Weiss & Oberw., 云南
59.Amanita pallidochlorotica Zhu L. Yang, Yang-Yang Cui & Qing Cai, 云南
60.Amanita pallidorosea P. Zhang & Zhu L. Yang, 重庆
61.Amanita pallidozonata Yang-Yang Cui, Qing Cai & Zhu L. Yang, 云南
62.Amanita pannosa Yang-Yang Cui, Qing Cai & Zhu L. Yang, 广东
63.Amanita parvicurta Yang-Yang Cui, Qing Cai & Zhu L. Yang, 云南
64.Amanita parviexitialis Qing Cai, Zhu L. Yang & Yang-Yang Cui, 广东
65.Amanita parvifritillaria Yang-Yang Cui, Qing Cai & Zhu L. Yang, 云南
66.Amanita parvipantherina Zhu L. Yang, M. Weiss & Oberw., 云南
67.Amanita parvivirginea Yang-Yang Cui, Qing Cai & Zhu L. Yang, 广东
68.Amanita pinophila Yang-Yang Cui, Qing Cai & Zhu L. Yang, 云南
69.Amanita pseudomanginiana Qing Cai, Yang-Yang Cui & Zhu L. Yang, 四川
70.Amanita pseudopantherina Zhu L. Yang ex Yang-Yang Cui, Qing Cai & Zhu L. Yang, 云南
71.Amanita pseudoprinceps Yang-Yang Cui, Qing Cai & Zhu L. Yang, 云南
72.Amanita pseudosychnopyramis Yang Y. Cui, Q. Cai & Zhu L Yang, 云南
73.Amanita pyramidata Zhu L. Yang, Yang-Yang Cui & Qing Cai, 云南
74.Amanita retenta Yang-Yang Cui, Qing Cai & Zhu L. Yang, 云南
75.Amanita rimosa P. Zhang & Zhu L. Yang, 湖南
76.Amanita roseolifolia Yang-Yang Cui, Qing Cai & Zhu L. Yang, 海南
77.Amanita rubiginosa Qing Cai, Yang-Yang Cui & Zhu L. Yang, 云南
78.Amanita rubroflava Yang-Yang Cui, Qing Cai & Zhu L. Yang, 云南
79.Amanita shennongjiana Yang-Yang Cui, Qing Cai & Zhu L. Yang, 湖北
80.Amanita sinensis Zhu L. Yang, 四川
81.Amanita sinensis var. subglobispora Zhu L. Yang, T.H. Li & X.L. Wu, 海南
82.Amanita sinocitrina Zhu L. Yang, Z.H. Chen & Z.G. Zhang, 湖南
83.Amanita squarrosipes Zhu L. Yang, Yang-Yang Cui & Qing Cai, 四川
84.Amanita subfrostiana Zhu L. Yang, 云南
85.Amanita subfuliginea Qing Cai, Zhu L. Yang & Yang-Yang Cui, 广东
86.Amanita subglobosa Zhu L. Yang, 四川
87.Amanita subhemibapha Zhu L. Yang, Yang-Yang Cui & Qing Cai, 云南
88.Amanita subparvipantherina Zhu L. Yang, Q. Cai & Yang Y. Cui, 云南
89.Amanita tenuifulva Yang-Yang Cui, Qing Cai & Zhu L. Yang, 云南
90.Amanita tomentosivolva Zhu L. Yang, 云南
91.Amanita verrucosivolva Zhu L. Yang, 云南
92.Amanita yenii Zhu L. Yang & C.M. Chen, 台湾
93.Amanita yuaniana Zhu L. Yang, 四川
94.Amanita zangii Zhu L. Yang, T.H. Li & X.L. Wu, 海南
95.Amanita zonata Yang-Yang Cui, Qing Cai & Zhu L. Yang, 云南
96.Amaropostia hainanensis B.K. Cui, L.L. Shen & Y.C. Dai, 海南
97.Amauroderma concentricum J. Song, Xiao L. He & B.K. Cui, 四川
98.Amyloporia pinea B.K. Cui & Y.C. Dai, 海南
99.Amylostereum orientale S.H. He & Hai J. Li, 安徽
100.Anamika angustilamellata Zhu L. Yang & Z.W. Ge, 云南
101.Anomoloma luteoalba J. Song & B.K. Cui, 浙江
102.Anomoloma rhizosum Y.C. Dai & Niemelä, 四川
103.Anomoloma submyceliosum J. Song & B.K. Cui, 福建
104.Anomoporia vesiculosa Y.C. Dai & Niemelä, 吉林
105.Antrodia bambusicola Y.C. Dai & B.K. Cui, 安徽
106.Antrodia hingganensis Y.C. Dai & Penttilä, 黑龙江
107.Antrodia huangshanensis Y.C. Dai & B.K. Cui, 安徽
108.Antrodia leucaena Y.C. Dai & Niemelä, 吉林
109.Antrodia monomitica Yuan Y. Chen, 黑龙江
110.Antrodia subserpens B.K. Cui & Yuan Y. Chen, 浙江
111.Antrodia subxantha Y.C. Dai & X.S. He, 四川
112.Antrodia tropica B.K. Cui, 海南
113.Antrodia wangii Y.C. Dai & H.S. Yuan, 北京
114.Antrodiella albocinnamomea Y.C. Dai & Niemelä, 吉林
115.Antrodiella chinensis H.S. Yuan, 吉林
116.Antrodiella citripileata H.S. Yuan, 云南
117.Antrodiella lactea H.S. Yuan, 广西
118.Antrodiella micra Y.C. Dai, 吉林
119.Antrodiella nanospora H.S. Yuan, 广西
120.Antrodiella pendulina H.S. Yuan, 广东
121.Antrodiella perennis B.K. Cui & Y.C. Dai, 江西
122.Antrodiella stipitata H.S. Yuan & Y.C. Dai, 黑龙江
123.Antrodiella thujae H.S. Yuan & Y.C. Dai, 青海
124.Antrodiella ussurii Y.C. Dai & Niemelä, 吉林
125.Aporpium miniporum F. Wu, Y.C. Dai & L.W. Zhou, 海南
126.Aureoboletus catenarius G. Wu & Zhu L. Yang, 云南
127.Aureoboletus nephrosporus G. Wu & Zhu L. Yang, 云南
128.Aureoboletus tomentosus G. Wu & Zhu L. Yang, 广东
129.Aureoboletus yunnanensis G. Wu & Zhu L. Yang, 云南
130.Auricularia heimuer F. Wu, B.K. Cui, Y.C. Dai, 黑龙江
131.Auricularia minutissima Y.C. Dai, F. Wu & Malysheva, 河北
132.Auricularia orientalis Y.C. Dai & F. Wu, 河北
133.Auricularia tibetica Y.C. Dai & F. Wu, 西藏
134.Auriscalpium microsporum P.M. Wang & Zhu L. Yang, 云南
135.Auriscalpium orientale P.M. Wang & Zhu L. Yang, 云南
136.Boidinia parva Ghobad-Nejhad, S.L. Liu, Y.C. Dai & E. Langer, 吉林
137.Boletellus indistinctus G. Wu, F. Li & Zhu L. Yang, 广东
138.Boletus botryoides B. Feng, Yang-Yang Cui, J.P. Xu & Zhu L. Yang, 湖南
139.Boletus fagacicola B. Feng, Yang-Yang Cui, J.P. Xu & Zhu L. Yang, 湖南
140.Boletus griseiceps B. Feng, Yang-Yang Cui, J.P. Xu & Zhu L. Yang, 福建
141.Boletus monilifer B. Feng, Yang-Yang Cui, J.P. Xu & Zhu L. Yang, 云南
142.Boletus orientialbus N.K. Zeng & Zhu L. Yang, 福建
143.Boletus sinoedulis B. Feng, Yang-Yang Cui, J.P. Xu & Zhu L. Yang, 四川
144.Boletus subviolaceofuscus B. Feng, Yang-Yang Cui, J.P. Xu & Zhu L. Yang, 云南
145.Boletus tylopilopsis B. Feng, Yang-Yang Cui, J.P. Xu & Zhu L. Yang, 云南
146.Boletus umbrinipileus B. Feng, Yang-Yang Cui, J.P. Xu & Zhu L. Yang, 云南
147.Boletus viscidiceps B. Feng, Yang-Yang Cui, J.P. Xu & Zhu L. Yang, 云南
148.Bondarzewia podocarpi Y.C. Dai & B.K. Cui, 海南
149.Bondarzewia submesenterica Jia J. Chen, B.K. Cui & Y.C. Dai, 云南
150.Bondarzewia tibetica B.K. Cui, J. Song & Jia J. Chen, 西藏
151.Bothia fujianensis N.K. Zeng & Zhu L. Yang, 福建
152.Buglossoporus eucalypticola M.L. Han, B.K. Cui & Y.C. Dai, 海南
153.Butyriboletus pseudospeciosus Kuan Zhao & Zhu L. Yang, 云南
154.Caloboletus yunnanensis Kuan Zhao and Zhu L. Yang, 云南
155.Cerarioporia cystidiata F. Wu, L.W. Zhou & J. Si, 海南
156.Ceriporia albomellea Yuan Yuan, Jia J. Chen & X.H. Ji, 海南
157.Ceriporia bubalinomarginata B.S. Jia & Y.C. Dai, 四川
158.Ceriporia crassitunicata Y.C. Dai & S.H. Wu, 台湾
159.Ceriporia inflata B.S. Jia & B.K. Cui, 海南
160.Ceriporia jiangxiensis B.S. Jia & B.K. Cui, 江西
161.Ceriporia nanlingensis B.K. Cui & B.S. Jia, 湖南
162.Ceriporia pseudocystidiata B.S. Jia & Y.C. Dai, 湖南
163.Ceriporia variegata B.S. Jia & Y.C. Dai, 湖南
164.Ceriporiopsis alboaurantia C.L. Zhao, B.K. Cui & Y.C. Dai, 福建
165.Ceriporiopsis egula C.J. Yu & Y.C. Dai, 西藏
166.Ceriporiopsis fimbriata C.L. Zhao & Y.C. Dai, 湖南
167.Ceriporiopsis lavendula B.K. Cui, 广东
168.Ceriporiopsis rosea C.L. Zhao & Y.C. Dai, 云南
169.Ceriporiopsis semisupina C.L. Zhao, B.K. Cui & Y.C. Dai, 浙江
170.Chalciporus rubinelloides G. Wu & Zhu L. Yang, 云南
171.Chiua angusticystidiata Yan C. Li & Zhu L. Yang, 云南
172.Chiua olivaceoreticulata Yan C. Li & Zhu L. Yang, 云南
173.Chiua viridula Yan C. Li & Zhu L. Yang, 云南
174.Chlorophyllum sphaerosporum Z.W. Ge & Zhu L. Yang, 内蒙古
175.Chroogomphus confusus Yan C. Li & Zhu L. Yang, 云南
176.Chroogomphus filiformis Yan C. Li & Zhu L. Yang, 云南
177.Chroogomphus orientirutilus Yan C. Li & Zhu L. Yang, 云南
178.Chroogomphus roseolus Yan C. Li & Zhu L. Yang, 云南
179.Cibaomyces glutinis Zhu L. Yang, Y.J. Hao & J. Qin, 云南
180.Climacocystis montana B.K. Cui & J. Song, 西藏
181.Clitocella orientalis S.P. Jian & Zhu L. Yang, 湖北
182.Clitopilopsis albida S.P. Jian & Zhu L. Yang, 云南
183.Clitopilus amygdaliformis Zhu L. Yang, 云南
184.Clitopilus brunneiceps S.P. Jian & Zhu L. Yang, 云南
185.Clitopilus rugosiceps S.P. Jian & Zhu L. Yang, 云南
186.Clitopilus sinoapalus S.P. Jian & Zhu L. Yang, 云南
187.Clitopilus umbilicatus S.P. Jian & Zhu L. Yang, 安徽
188.Clitopilus yunnanensis S.P. Jian & Zhu L. Yang, 云南
189.Coltricia abieticola Y.C. Dai, 云南
190.Coltricia austrosinensis L.S. Bian & Y.C. Dai, 云南
191.Coltricia crassa Y.C. Dai, 云南
192.Coltricia lateralis L.S. Bian & Y.C. Dai, 云南
193.Coltricia macropora Y.C. Dai, 广东
194.Coltricia minima L.S. Bian & Y.C. Dai, 海南
195.Coltricia minor Y.C. Dai, 湖南
196.Coltricia rigida L.S. Bian & Y.C. Dai, 海南
197.Coltricia spina Y.C. Dai, 湖南
198.Coltricia subcinnamomea L.S. Bian & Y.C. Dai, 山西
199.Coltricia subperennis Y.C. Dai, 海南
200.Coltricia subverrucata L.S. Bian & Y.C. Dai, 贵州
201.Coltricia tsugicola Y.C. Dai & B.K. Cui, 福建
202.Coltricia weii Y.C. Dai, 湖南
203.Coltricia wenshanensis L.S. Bian & Y.C. Dai, 云南
204.Coltriciella baoshanensis Y.C. Dai & B.K. Cui, 云南
205.Coltriciella globosa L.S. Bian & Y.C. Dai, 广东
206.Coltriciella naviculiformis Y.C. Dai & Niemelä, 云南
207.Coltriciella pseudodependens L.S. Bian & Y.C. Dai, 云南
208.Coltriciella subglobosa Y.C. Dai, 海南
209.Cordyceps chaetoclavata H. Yu, Y.B. Wang, Y. Wang, Q. Fan & Zhu L. Yang, 云南
210.Cordyceps cocoonihabita H. Yu, Y.B. Wang, Y. Wang, Q. Fan & Zhu L. Yang, 云南
211.Cordyceps shuifuensis H. Yu, Y.B. Wang, Y. Wang & Zhu L. Yang, 云南
212.Cordyceps subtenuipes H. Yu, Y.B. Wang, Y. Wang, D.E. Duan & Zhu L. Yang, 云南
213.Coriolopsis dendriformis Hai J. Li, Y.C. Dai & B.K. Cui, 海南
214.Coriolopsis hainanensis Hai J. Li, Y.C. Dai & B.K. Cui, 海南
215.Crassisporus imbricatus B.K. Cui & Xing Ji, 海南
216.Crassisporus macroporus B.K. Cui & Xing Ji, 广西
217.Crassisporus microsporus B.K. Cui & Xing Ji, 云南
218.Cyanoboletus brunneoruber G. Wu & Zhu L. Yang, 云南
219.Cyanosporus fusiformis B.K. Cui, L.L. Shen & Y.C. Dai, 贵州
220.Cyanosporus microporus B.K. Cui, L.L. Shen & Y.C. Dai, 云南
221.Cyanosporus mongolicus B.K. Cui, L.L. Shen & Y.C. Dai, 内蒙古
222.Cyanosporus piceicola B.K. Cui, L.L. Shen & Y.C. Dai, 四川
223.Cyanosporus subhirsutus B.K. Cui, L.L. Shen & Y.C. Dai, 贵州
224.Cyanosporus tricolor B.K. Cui, L.L. Shen & Y.C. Dai, 西藏
225.Cyanosporus ungulatus B.K. Cui, L.L. Shen & Y.C. Dai, 四川
226.Cyclomyces lamellatus Y.C. Dai & Niemelä, 湖南
227.Cyphellopsis changbaiensis Y.L. Wei & W.M. Qin, 吉林
228.Cyptotrama glabra Zhu L. Yang & J. Qin, 云南
229.Cyptotrama shennongjia Zhu L. Yang & J. Qin, 湖北
230.Dacryobolus angiospermarum S.H. He, 福建
231.Dacryobolus gracilis H.S. Yuan, 重庆
232.Dacryobolus montanus X.Z. Wan & H.S. Yuan, 广西
233.Dacryoscyphus chrysochilus R. Kirschner & Zhu L. Yang, 云南
234.Daedalea allantoidea M.L. Han, B.K. Cui & Y.C. Dai, 云南
235.Daedalea circularis B.K. Cui & Hai J. Li, 云南
236.Daedalea radiata B.K. Cui & Hai J. Li, 云南
237.Daedaleopsis hainanensis Hai J. Li & S.H. He, 海南
238.Datroniella melanocarpa B.K. Cui, Hai J. Li & Y.C. Dai, 四川
239.Datroniella subtropica B.K. Cui, Hai J. Li & Y.C. Dai, 四川
240.Datroniella tibetica B.K. Cui, Hai J. Li & Y.C. Dai, 西藏
241.Datroniella tropica B.K. Cui, Hai J. Li & Y.C. Dai, 云南
242.Dendrodontia hyphopaxillosa M.J. Li & H.S. Yuan, 广西
243.Dendrodontia taiwaniana H.C. Wang & Sheng H. Wu, 台湾
244.Dentipellis coniferarum Y.C. Dai & L.W. Zhou, 吉林
245.Dentipellis microspora Y.C. Dai, 吉林
246.Dentipellis rhizomorpha Yuan Yuan & Y.C. Dai, 云南
247.Dentipellopsis dacrydicola Y.C. Dai & L.W. Zhou, 海南
248.Dextrinocystis calamicola S.H. He & S.L. Liu, 福建
249.Dextrinoporus aquaticus H.S. Yuan, gen. & sp. nov., 广西
250.Dichomitus hubeiensis Hai J. Li & B.K. Cui, 湖北
251.Dichomitus sinuolatus H.S. Yuan, 广东
252.Dichostereum austrosinense S.H. He & S.L. Liu, 广西
253.Dichostereum boidinii S.H. He & S.L. Liu, 湖北
254.Dichostereum eburneum S.H. He & S.L. Liu, 福建
255.Donkioporiella mellea L.W. Zhou, 广西
256.Elmerina fragilis F. Wu & Hai J. Li, 河南
257.Epithele cylindricosterigmata H.C. Wang & Sheng H. Wu, 台湾
258.Epithele lutea H.C. Wang & Sheng H. Wu, 台湾
259.Exidia yadongensis F. Wu, Qi Zhao, Zhu L. Yang & Y.C. Dai, 西藏
260.Favolus niveus J.L. Zhou & B.K. Cui, 云南
261.Favolus pseudoemerici J.L. Zhou & B.K. Cui, 海南
262.Favolus septatus J.L. Zhou & B.K. Cui, 广西
263.Favolus subtropicus J.L. Zhou & B.K. Cui, 福建
264.Fibroporia albicans B.K. Cui & Yuan Y. Chen, 江西
265.Fibroporia bambusae Yuan Y. Chen & B.K. Cui, 海南
266.Fibroporia ceracea Yuan Y. Chen & B.K. Cui, 云南
267.Fistulina subhepatica B.K. Cui & J. Song, 云南
268.Flammeopellis bambusicola Y.C. Dai, B.K. Cui & C.L. Zhao, 四川
269.Flammulina yunnanensis Z.W. Ge & Zhu L. Yang, 云南
270.Fomitiporella austroasiana Y.C. Dai, X.H. Ji & J. Vlasák, 海南
271.Fomitiporella caviphila L.W. Zhou, 浙江
272.Fomitiporella sinica Y.C. Dai, X.H. Ji & Vlasák, 江西
273.Fomitiporella subinermis Y.C. Dai, X.H. Ji & Vlasák, 湖南
274.Fomitiporia alpina B.K. Cui & Hong Chen, 四川
275.Fomitiporia bannaensis Y.C. Dai, 云南
276.Fomitiporia ellipsoidea B.K. Cui & Y.C. Dai, 福建
277.Fomitiporia gaoligongensis B.K. Cui & Hong Chen, 云南
278.Fomitiporia hainaniana B.K. Cui & Hong Chen, 海南
279.Fomitiporia norbulingka B.K. Cui & Hong Chen, 西藏
280.Fomitiporia pentaphylacis L.W. Zhou, 广西
281.Fomitiporia punicata Y.C. Dai, B.K. Cui & Decock, 陕西
282.Fomitiporia rhamnoides T.Z. Liu & F. Wu, 河北
283.Fomitiporia subhippophaëicola B.K. Cui & H. Chen, 西藏
284.Fomitiporia subrobusta B.K. Cui & Hong Chen, 云南
285.Fomitiporia subtropica B.K. Cui & Hong Chen, 广东
286.Fomitiporia tenuitubus L.W. Zhou, 广西
287.Fomitiporia tibetica Y.C. Dai & M. Zang, 西藏
288.Fomitiporia torreyae Y.C. Dai & B.K. Cui, 福建
289.Fomitopsis cana B.K. Cui, Hai J. Li & M.L. Han, 海南
290.Fomitopsis cystidiata B.K. Cui & M.L. Han, 海南
291.Fomitopsis fragilis B.K. Cui & M.L. Han, 海南
292.Fomitopsis ginkgonis B.K. Cui & Shun Liu, 湖北
293.Fomitopsis niveomarginata L.W. Zhou & Y.L. Wei, 吉林
294.Fomitopsis subfeei B.K. Cui & M.L. Han, 四川
295.Fomitopsis subtropica B.K. Cui, Hai J. Li & M.L. Han, 广东
296.Foraminispora yinggelingensis Y.F. Sun & B.K. Cui, 海南
297.Fragiliporia fragilis Y.C. Dai, B.K. Cui & C.L. Zhao, 云南
298.Frantisekia abieticola H.S. Yuan, 云南
299.Fulvifomes hainanensis L.W. Zhou, 海南
300.Fulvoderma australe L.W. Zhou & Y.C. Dai, 湖南
301.Funalia cystidiata Hai J. Li, Y.C. Dai & B.K. Cui, 海南
302.Funalia subgallica Hai J. Li & S.H. He, 海南
303.Fuscoporia acutimarginata Y.C. Dai & Q. Chen, 云南
304.Fuscoporia australasica Q. Chen, F. Wu & Y.C. Dai, 云南
305.Fuscoporia chinensis Q. Chen, F. Wu & Y.C. Dai, 云南
306.Fuscoporia karsteniana Q. Chen, F. Wu & Y.C. Dai, 吉林
307.Fuscoporia montana Y.C. Dai & Niemelä, 云南
308.Fuscoporia monticola Y.C. Dai, Q. Chen & J. Vlasák, 安徽
309.Fuscoporia ramulicola Y.C. Dai & Q. Chen, 云南
310.Fuscoporia sinica Y.C. Dai, Q. Chen & J. Vlasák, 宁夏
311.Fuscoporia subchrysea Q. Chen, F. Wu & Y.C. Dai, 海南
312.Fuscoporia subferrea Q. Chen & Yuan Yuan, 海南
313.Fuscoporia yunnanensis Y.C. Dai, 云南
314.Ganoderma angustisporum J.H. Xing, B.K. Cui & Y.C. Dai, 福建
315.Ganoderma casuarinicola J.H. Xing, B.K. Cui & Y.C. Dai, 广东
316.Ganoderma lingzhi Sheng H. Wu, Y. Cao & Y.C. Dai, 湖北
317.Ganoderma mutabile Y. Cao & H.S. Yuan, 云南
318.Geliporus exilisporus (Y.C. Dai & Niemelä) Yuan Yuan, Jia J. Chen & S.H. He, 广东
319.Gloeodontia eriobotryae Y.C. Dai & L.W. Zhou, 海南
320.Gloeoporus citrinoalbus Yuan Yuan & Jia J. Chen, 海南
321.Gloeoporus hainanensis Yuan Yuan & Jia J. Chen, 海南
322.Gloiocephala parvinelumbonifolia Chun Y. Deng, J. Qin & Zhu L. Yang, 海南
323.Grammatus labyrinthinus H.S. Yuan & C. Decock, 云南
324.Grammothele denticulata Y.C. Dai & L.W. Zhou, 广东
325.Grammothele hainanensis F. Wu & L.W. Zhou, 海南
326.Grammothele separabillima H.S. Yuan, 广西
327.Grammothelopsis asiatica Y.C. Dai & B.K. Cui, 云南
328.Grammothelopsis subtropica B.K. Cui & C.L. Zhao, 广东
329.Griseoporia taiwanense Y.C. Dai & S.H. He, 台湾
330.Gyromitra venenata Hai J. Li, Z.H. Chen & Zhu L.Yang, 湖南
331.Hapalopilus flavus B.K. Cui & Y.C. Dai, 福建
332.Haploporus angustisporus Meng Zhou & Y.C. Dai, 广东
333.Haploporus crassus Meng Zhou & Y.C. Dai, 云南
334.Haploporus cylindrosporus L.L. Shen, Y.C. Dai & B.K. Cui, 云南
335.Haploporus latisporus Juan Li & Y.C. Dai, 河南
336.Haploporus microsporus L.L. Shen, Y.C. Dai & B.K. Cui, 海南
337.Haploporus septatus L.L. Shen, Y.C. Dai & B.K. Cui, 云南
338.Haploporus subpapyraceus L.L. Shen, Y.C. Dai & B.K. Cui, 浙江
339.Harrya alpina Yan C. Li & Zhu L. Yang, 云南
340.Harrya atrogrisea Yan C. Li & Zhu L. Yang, 云南
341.Harrya moniliformis Yan C. Li & Zhu L. Yang, 云南
342.Harrya subalpina Yan C. Li & Zhu L. Yang, 云南
343.Heimioporus conicus N.K. Zeng & Zhu L. Yang, 海南
344.Heimioporus gaojiaocong N.K. Zeng & Zhu L. Yang, 云南
345.Helvella bachu Q. Zhao, Zhu L. Yang & K.D. Hyde, 新疆
346.Helvella involuta Q. Zhao, Zhu L. Yang & K.D. Hyde, 云南
347.Helvella orienticrispa Q. Zhao, Zhu L. Yang & K.D. Hyde, 云南
348.Helvella pseudoreflexa Q. Zhao, Zhu L. Yang & K.D. Hyde, 云南
349.Helvella subspadicea Q. Zhao, Zhu L. Yang & K.D. Hyde, 新疆
350.Hemileccinum rugosum G. Wu & Zhu L. Yang, 新疆
351.Henningsomyces leptus Y.L. Wei & Y.C. Dai, 吉林
352.Henningsomyces subiculatus Y.L. Wei & W.M. Qin, 广西
353.Heterobasidion amyloideum Y.C. Dai, Jia J. Chen & Korhonen, 西藏
354.Heterobasidion armandii Y.C. Dai, Jia J. Chen & Yuan Yuan, 云南
355.Heterobasidion australe Y.C. Dai & Korhonen, 江西
356.Heterobasidion linzhiense Y.C. Dai & Korhonen, 西藏
357.Heterobasidion subinsulare Y.C. Dai, Jia J. Chen & Yuan Yuan, 云南
358.Heterobasidion subparviporum Y.C. Dai, Jia J. Chen & Yuan Yuan, 河北
359.Heterobasidion tibeticum Y.C. Dai, Jia J. Chen & Korhonen, 西藏
360.Hortiboletus amygdalinus Xue T. Zhu & Zhu L. Yang, 云南
361.Hydnochaete asetosa Y.C. Dai, 海南
362.Hydnocristella latihypha Jia J. Chen, L.L. Shen & B.K. Cui, 四川
363.Hymenagaricus taiwanensis Z.W. Ge, Zhu L. Yang & C.M. Chen, 台湾
364.Hymenoboletus luteopurpureus Yan C. Li & Zhu L. Yang, 云南
365.Hymenochaete acerosa S.H. He & Hai J. Li, 西藏
366.Hymenochaete angustispora S.H. He & Y.C. Dai, 山西
367.Hymenochaete biformisetosa Jiao Yang & S.H. He, 云南
368.Hymenochaete cana S.H. He & Hai J. Li, 广西
369.Hymenochaete fissurata S.H. He & Hai J. Li, 云南
370.Hymenochaete huangshanensis S.H. He & Y.C. Dai, 安徽
371.Hymenochaete macrospora Y.C. Dai, 吉林
372.Hymenochaete megaspora S.H. He & Hai J. Li, 西藏
373.Hymenochaete micropora L.W. Zhou & Y.C. Dai, 云南
374.Hymenochaete minor S.H. He & Y.C. Dai, 广西
375.Hymenochaete orientalis S.H. He, 台湾
376.Hymenochaete parmastoi S.H. He & Hai J. Li, 西藏
377.Hymenochaete quercicola S.H. He & Hai J. Li, 西藏
378.Hymenochaete rhododendricola S.H. He & Hai J. Li, 西藏
379.Hymenochaete subporioides L.W. Zhou & Y.C. Dai, 浙江
380.Hymenochaete tropica S.H. He & Y.C. Dai, 云南
381.Hymenochaete verruculosa S.H. He & Y.C. Dai, 山西
382.Hymenochaete yunnanensis S.H. He & Hai J. Li, 云南
383.Hymenochaetopsis laricicola S.H. He & Jiao Yang, 黑龙江
384.Hyphodermella poroides Y.C. Dai & C.L. Zhao, 海南
385.Hyphodontia bubalina Min Wang, Yuan Y. Chen & B.K. Cui, 内蒙古
386.Hyphodontia capitatocystidiata H.X. Xiong, Y.C. Dai & Sheng H. Wu, 台湾
387.Hyphodontia dimitica Jia J. Chen & L.W. Zhou, 湖南
388.Hyphodontia heterocystidiata H.X. Xiong, Y.C. Dai & Sheng H. Wu, 台湾
389.Hyphodontia mollissima L.W. Zhou, 海南
390.Hyphodontia mongolica Min Wang, Yuan Y. Chen & B.K. Cui, 内蒙古
391.Hyphodontia pseudotropica C.L. Zhao, B.K. Cui & Y.C. Dai, 海南
392.Hyphodontia rhizomorpha C.L. Zhao, B.K. Cui & Y.C. Dai, 云南
393.Hyphodontia septocystidiata H.X. Xiong, Y.C. Dai & Sheng H. Wu, 台湾
394.Hyphodontia sinensis H.X. Xiong, Y.C. Dai & Sheng H. Wu, 黑龙江
395.Hyphodontia subefibulata Jia J. Chen & L.W. Zhou, 江西
396.Hyphodontia subpallidula H.X. Xiong, Y.C. Dai & Sheng H. Wu, 台湾
397.Hyphodontia syringae Langer, 吉林
398.Hypochnicium austrosinensis W.M. Qin & L.W. Zhou, 湖南
399.Imleria parva Xue T. Zhu & Zhu L. Yang, 云南
400.Imleria subalpina Xue T. Zhu & Zhu L. Yang, 云南
401.Inocutis subdryophila Y.C. Dai & H.S. Yuan, 西藏
402.Inonotus acutus B.K. Cui & Y.C. Dai, 海南
403.Inonotus alpinus Y.C. Dai & X.M. Tian, 西藏
404.Inonotus canaricola Y.C. Dai, 海南
405.Inonotus castanopsidis G.J. Ren & Y.C. Dai, 云南
406.Inonotus casuarinae L.S. Bian, 海南
407.Inonotus chrysomarginatus B.K. Cui & Y.C. Dai, 海南
408.Inonotus exilisporus Y.C. Dai & Niemelä, 辽宁
409.Inonotus griseus L.W. Zhou, 安徽
410.Inonotus hainanensis Y.C. Dai & Xiong, 海南
411.Inonotus henanensis Juan Li & Y.C. Dai, 河南
412.Inonotus indurescens Y.C. Dai, 云南
413.Inonotus latemarginatus Y.C. Dai, 海南
414.Inonotus magnisetus Y.C. Dai, 广东
415.Inonotus niveomarginatus H.Y. Yu, C.L. Zhao & Y.C. Dai, 云南
416.Inonotus puerensis Hai J. Li & S.H. He, 云南
417.Inonotus rigidus B.K. Cui & Y.C. Dai, 云南
418.Inonotus sanghuang Sheng H. Wu, T. Hatt. & Y.C. Dai, 吉林
419.Inonotus sublevis Y.C. Dai & Niemelä, 云南
420.Inonotus tenuicontextus L.W. Zhou & W.M. Qin, 贵州
421.Inonotus tenuissimus H.Y. Yu, C.L. Zhao & Y.C. Dai, 云南
422.Inonotus zonatus Y.C. Dai & X.M. Tian, 海南
423.Junghuhnia austrosinensis F. Wu, P. Du & X.M. Tian, 云南
424.Junghuhnia flabellata H.S. Yuan & Y.C. Dai, 广西
425.Junghuhnia minor H.S. Yuan, 云南
426.Junghuhnia nandinae F. Wu, P. Du & X.M. Tian, 重庆
427.Junghuhnia rhizomorpha H.S. Yuan & Y.C. Dai, 云南
428.Junghuhnia subcollabens F. Wu, P. Du & X.M. Tian, 云南
429.Junghuhnia subnitida H.S. Yuan & Y.C. Dai, 云南
430.Junghuhnia pseudocrustacea H.S. Yuan, 广西
431.Junghuhnia pseudominuta H.S. Yuan & Y.C. Dai, 海南
432.Junghuhnia rhizomorpha H.S. Yuan & Y.C. Dai, 云南
433.Junghuhnia subnitida H.S. Yuan & Y.C. Dai, 云南
434.Junghuhnia taiwaniana H.S. Yuan, Sheng H. Wu & Y.C. Dai, 台湾
435.Junghuhnia tropica H.S. Yuan, Sheng H. Wu & Y.C. Dai, 台湾
436.Laccaria alba Zhu L. Yang & Lan Wang, 云南
437.Laccaria angustilamella Zhu L. Yang & Lan Wang, 云南
438.Laccaria moshuijun Popa & Zhu L. Yang, 云南
439.Laetiporus ailaoshanensis B.K. Cui & J. Song, 云南
440.Laetiporus medogensis J. Song & B.K. Cui, 西藏
441.Laetiporus xinjiangensis J. Song, Y.C. Dai & B.K. Cui, 新疆
442.Laetiporus zonatus B.K. Cui & J. Song, 云南
443.Lanmaoa angustispora G. Wu & Zhu L. Yang, 云南
444.Lanmaoa asiatica G. Wu & Zhu L. Yang, 云南
445.Larssoniporia incrustatocystidiata Y.C. Dai, Jia J. Chen & B.K. Cui, 云南
446.Lauriliella taiwanensis S.H. He & Nakasone, 台湾
447.Leccinellum cremeum Zhu L. Yang & G. Wu, 云南
448.Leifia brevispora Gafforov, S.L. Liu & L.W. Zhou, 湖北
449.Leifiporia rhizomorpha Y.C. Dai, F. Wu & C.L. Zhao, 安徽
450.Lenzitopsis daii L.W. Zhou & Kõljalg, 四川
451.Lepiota angusticystidiata J.F. Liang & Z.L. Yang, 云南
452.Lepiota attenuata J.F. Liang & Zhu L. Yang, 云南
453.Lepiota brunneosquamulosa J.F. Liang & Z.L. Yang, 云南
454.Lepiota catenariocystidia H.C. Wang & Zhu L. Yang, 云南
455.Lepiota cristatanea J.F. Liang & Zhu L. Yang, 云南
456.Lepiota furfuraceipes H.C. Wang & Zhu L. Yang, 云南
457.Lepiota venenata Z.H. Chen & Zhu L. Yang, 湖北
458.Leucoagaricus atroazureus J.F. Liang, Zhu L. Yang & J. Xu, 云南
459.Leucoagaricus flavovirens J.F. Liang, Zhu L. Yang & J. Xu, 海南
460.Leucoagaricus subcrystallifer Z.W. Ge & Zhu L. Yang, 四川
461.Leucoagaricus subpurpureolilacinus Z.W. Ge & Zhu L. Yang, 云南
462.Leucoagaricus truncatus Z.W. Ge & Zhu L. Yang, 四川
463.Liangia sinensis H. Yu, Y.B. Wang, Y. Wang, Z.H. Chen & Zhu L. Yang, 云南
464.Lignosus hainanensis B.K. Cui, 海南
465.Limacella taiwanensis Zhu L. Yang & W.N. Chou, 台湾
466.Limacellopsis asiatica Zhu L. Yand, Qing Cai & Yang-Yang Cui, 四川
467.Lindtneria asiae-orientalis S.L. Liu & S.H. He, 黑龙江
468.Lopharia resupinata S.H. He, S.L. Liu & Y.C. Dai, 江西
469.Lopharia sinensis S.H. He, S.L. Liu & Y.C. Dai, 宁夏
470.Luteoporia albomarginata F. Wu, Jia J. Chen & S.H. He, 海南
471.Macrolepiota detersa Z.W. Ge, Zhu L. Yang & Vellinga, 安徽
472.Macrolepiota orientiexcoriata Z.W. Ge, Zhu L. Yang & Vellinga, 四川
473.Macrolepiota velosa Vellinga & Zhu L. Yang, 云南
474.Megasporia guangdongensis B.K. Cui & Hai J. Li, 广东
475.Megasporia hengduanensis B.K. Cui & Hai J. Li, 云南
476.Megasporia rimosa Y. Yuan, X.H. Ji & Y.C. Dai, 广西
477.Megasporia tropica Y. Yuan, X.H. Ji & Y.C. Dai, 海南
478.Megasporia yunnanensis Y. Yuan, X.H. Ji & Y.C. Dai, 云南
479.Megasporoporia bannaensis B.K. Cui & Hai J. Li, 云南
480.Megasporoporia cystidiolophora B.K. Cui & Y.C. Dai, 浙江
481.Megasporoporia ellipsoidea B.K. Cui & P. Du, 海南
482.Megasporoporia minor B.K. Cui & Hai J. Li, 云南
483.Megasporoporia minuta Y.C. Dai & X.S. Zhou, 广西
484.Megasporoporia quercina Y.C. Dai, 云南
485.Megasporoporia rhododendri Y.C. Dai & Y.L. Wei, 四川
486.Megasporoporia subcavernulosa Y.C. Dai & Sheng H. Wu, 云南
487.Megasporoporia violacea B.K. Cui & P. Du, 海南
488.Megasporoporiella lacerata B.K. Cui & Hai J. Li, 云南
489.Megasporoporiella pseudocavernulosa B.K. Cui & Hai J. Li, 云南
490.Melanoderma disciforme H.S. Yuan, 云南
491.Melanoderma microcarpum B.K. Cui & Y.C. Dai, 湖南
492.Mensularia lithocarpi L.W. Zhou, 云南
493.Mensularia rhododendri F. Wu, Y.C Dai & L.W. Zhou, 贵州
494.Michenera incrustata S.H. He, S.L. Liu & Nakasone, 云南
495.Mycoacia angustata H.S. Yuan, 海南
496.Mycoleptodonoides tropicalis H.S. Yuan & Y.C. Dai, 云南
497.Mycorrhaphium sessile H.S. Yuan & Y.C. Dai, 云南
498.Myxoderma olivaceum Zhu L. Yang, Yang-Yang Cui & Qing Cai, 云南
499.Neoalbatrellus odorus Yuan Y. Chen & B.K. Cui, 云南
500.Neodatronia gaoligongensis B.K. Cui, Hai J. Li & Y.C. Dai, 云南
501.Neodatronia sinensis B.K. Cui, Hai J. Li & Y.C. Dai, 安徽
502.Neofavolus squamatus J.H. Xing, J.L. Zhou & B.K. Cui, 西藏
503.Neofomitella guangxiensis B.K. Cui & Xing Ji, 广西
504.Neofomitella polyzonata Y.C. Dai, Hai J. Li & Vlasák, 江西
505.Neomensularia duplicata F. Wu, L.W. Zhou & Y.C. Dai, 海南
506.Neomensularia rectiseta X.H. Ji, L.W. Zhou & F. Wu, 湖南
507.Nigrofomes sinomelanoporus L.W. Zhou, 海南
508.Nigrofomes submelanoporus Meng Zhou & F. Wu, 云南
509.Odontia aculeata Yuan Yuan, Y.C. Dai & H.S. Yuan, 辽宁
510.Odontia sparsa Yuan Yuan, Y.C. Dai & H.S. Yuan, 辽宁
511.Omphalotus flagelliformis Zhu L. Yang & B. Feng, 云南
512.Onnia microspora Y.C. Dai & L.W. Zhou, 安徽
513.Onnia tibetica Y.C. Dai & S.H. He, 西藏
514.Ophiocordyceps highlandensis Zhu L. Yang & J. Qin, 云南
515.Otidea bicolor W.Y. Zhuang & Zhu L. Yang, 云南
516.Oudemansiella amygdaliformis Zhu L. Yang & M. Zang, 云南
517.Oudemansiella bii Zhu L. Yang & L.F. Zhang, 广东
518.Oudemansiella orientalis Zhu L. Yang, 云南
519.Oudemansiella velata Zhu L. Yang & M. Zang, 西藏
520.Oudemansiella yunnanensis Zhu L. Yang & M. Zang, 云南
521.Ovipoculum album Zhu L. Yang & R. Kirschner, 云南
522.Oxyporus ginkgonis Y.C. Dai, 北京
523.Oxyporus macroporus Y.C. Dai & Y.L. Wei, 四川
524.Oxyporus piceicola B.K. Cui & Y.C. Dai, 青海
525.Oxyporus subpopulinus B.K. Cui & Y.C. Dai, 青海
526.Panellus edulis Y.C. Dai & Niemelä, 吉林
527.Paraxerula ellipsospora Zhu L. Yang & J. Qin, 云南
528.Parvixerocomus pseudoaokii G. Wu, K. Zhao & Zhu L. Yang, 广东
529.Perenniporia aridula B.K. Cui & C.L. Zhao, 云南
530.Perenniporia bannaensis B.K. Cui & C.L. Zhao, 云南
531.Perenniporia cinereofusca B.K. Cui & C.L. Zhao, 海南
532.Perenniporia citrinoalba B.K. Cui, C.L. Zhao & Y.C. Dai, 海南
533.Perenniporia cystidiata Y.C. Dai & S.H. Wu, 台湾
534.Perenniporia hainaniana B.K. Cui & C.L. Zhao, 海南
535.Perenniporia hattorii Y.C. Dai & B.K. Cui, 海南
536.Perenniporia lacerata B.K. Cui & C.L. Zhao, 河南
537.Perenniporia luteola B.K. Cui & C.L. Zhao, 湖南
538.Perenniporia macropora B.K. Cui & C.L. Zhao, 广西
539.Perenniporia minor Y.C. Dai & H.X. Xiong, 吉林
540.Perenniporia nanlingensis B.K. Cui & C.L. Zhao, 广东
541.Perenniporia piceicola Y.C. Dai, 云南
542.Perenniporia punctata Hai J. Li & Jing Si, 湖北
543.Perenniporia pyricola Y.C. Dai & B.K. Cui, 辽宁
544.Perenniporia rhizomorpha B.K. Cui, Y.C. Dai & Decock, 安徽
545.Perenniporia russeimarginata B.K. Cui & C.L. Zhao, 云南
546.Perenniporia subcorticola Chao G. Wang & F. Wu, 福建
547.Perenniporia substraminea B.K. Cui & C.L. Zhao, 浙江
548.Perenniporia subtephropora B.K. Cui & C.L. Zhao, 广东
549.Perenniporia tianmuensis B.K. Cui & C.L. Zhao, 浙江
550.Perenniporia tibetica B.K. Cui & C.L. Zhao, 西藏
551.Perenniporia yinggelingensis B.K. Cui & Y.C. Dai, 海南
552.Phaeotremella yunnanensis L.F. Fan, F. Wu & Y.C. Dai, 云南
553.Phallus dongsun T.H. Li, T. Li, Chun Y. Deng, W.Q. Deng & Zhu L. Yang, 贵州
554.Phanerochaete aurantiobadia Ghobad- Nejhad, S.L. Liu & E. Langer, 吉林
555.Phanerochaete bambucicola Sheng H. Wu, 台湾
556.Phanerochaete cinerea Y.L. Xu & S.H. He, 海南
557.Phanerochaete hainanensis S.H. He & Y.C. Dai, 海南
558.Phanerochaete hymenochaetoides Y.L. Xu & S.H. He, 海南
559.Phanerochaete incarnata Sheng H. Wu, 台湾
560.Phanerochaete leptocystidiata Y.L. Xu & S.H. He, 广东
561.Phanerochaete metuloidea Y.L. Xu & S.H. He, 云南
562.Phanerochaete minor Y.L. Xu & S.H. He, 海南
563.Phanerochaete porostereoides S.L. Liu & S.H. He, 陕西
564.Phanerochaete sinensis Y.L. Xu, C.C. Chen & S.H. He, 辽宁
565.Phanerochaete subrosea Y.L. Xu & S.H. He, 宁夏
566.Phanerochaete yunnanensis Y.L. Xu & S.H. He, 云南
567.Phellinidium aciferum Y.C. Dai, 吉林
568.Phellinidium asiaticum Spirin, L.W. Zhou & Y.C. Dai, 吉林
569.Phellinidium qilianense B.K. Cui, L.W. Zhou & Y.C. Dai, 青海
570.Phellinopsis asetosa L.W. Zhou, 云南
571.Phellinopsis helwingiae W.M. Qin & L.W. Zhou, 四川
572.Phellinopsis junipericola L.W. Zhou, 青海
573.Phellinopsis lonicericola L.W. Zhou, 西藏
574.Phellinopsis resupinata L.W. Zhou, 河南
575.Phellinopsis tibetica L.W. Zhou, 西藏
576.Phellinus austrosinensis L.S. Bian, 海南
577.Phellinus bambusicola L.W. Zhou & B.S. Jia, 海南
578.Phellinus castanopsidis B.K. Cui, Y.C. Dai & Decock, 广东
579.Phellinus collinus Y.C. Dai & Niemelä, 湖南
580.Phellinus himalayensis Y.C. Dai, 云南
581.Phellinus minisporus B.K. Cui & Y.C. Dai, 海南
582.Phellinus monticola L.W. Zhou & Y.C. Dai, 西藏
583.Phellinus mori Y.C. Dai & B.K. Cui, 黑龙江
584.Phellinus orientoasiaticus L.W. Zhou & Y.C. Dai, 西藏
585.Phellinus padicola L.W. Zhou & Y.C. Dai, 青海
586.Phellinus parmastoi L.W. Zhou & Y.C. Dai, 吉林
587.Phellinus piceicola B.K. Cui & Y.C. Dai, 云南
588.Phellinus pseudoigniarius Y.C. Dai & Fan Yang, 新疆
589.Phellinus tenuiculus B.K. Cui, 江西
590.Phellodon subconfluens H.S. Yuan & F. Wu, 辽宁
591.Phlebiporia bubalina Jia J. Chen, B.K. Cui & Y.C. Dai, 云南
592.Phylloporia clausenae L.W. Zhou, 海南
593.Phylloporia crataegi L.W. Zhou & Y.C. Dai, 辽宁
594.Phylloporia cylindrispora L.W. Zhou, 广西
595.Phylloporia cystidiolophora F. Wu, G.J. Ren & Y.C. Dai, 重庆
596.Phylloporia dependens Y.C. Dai, 云南
597.Phylloporia flacourtiae L.W. Zhou, 广西
598.Phylloporia fontanesiae L.W. Zhou & Y.C. Dai, 河南
599.Phylloporia gutta L.W. Zhou & Y.C. Dai, 四川
600.Phylloporia hainaniana Y.C. Dai & B.K. Cui, 海南
601.Phylloporia homocarnica L.W. Zhou, 广西
602.Phylloporia lespedezae G.J. Ren & F. Wu, 山西
603.Phylloporia manglietiae Yuan Y. Chen & B.K. Cui, 海南
604.Phylloporia minutipora L.W. Zhou, 海南
605.Phylloporia nandinae L.W. Zhou & Y.C. Dai, 江西
606.Phylloporia oblongospora Y.C. Dai & H.S. Yuan, 广西
607.Phylloporia oreophila L.W. Zhou & Y.C. Dai, 西藏
608.Phylloporia osmanthi L.W. Zhou, 广西
609.Phylloporia pendula Yuan Y. Chen & B.K. Cui, 海南
610.Phylloporia perangusta F. Wu, G.J. Ren & Y.C. Dai, 广东
611.Phylloporia pseudopectinata Yuan Y. Chen & B.K. Cui, 海南
612.Phylloporia radiata L.W. Zhou, 贵州
613.Phylloporia rattanicola F. Wu, G.J. Ren & Y.C. Dai, 福建
614.Phylloporia splendida F. Wu, G.J. Ren & Y.C. Dai, 云南
615.Phylloporia subpulla F. Wu, G.J. Ren & Y.C. Dai, 海南
616.Phylloporia terrestris L.W. Zhou, 广西
617.Phylloporia tiliae L.W. Zhou, 湖南
618.Physisporinus crataegi F. Wu, Jia J. Chen & Y.C. Dai, 天津
619.Physisporinus lavendulus F. Wu, Jia J. Chen & Y.C. Dai, 海南
620.Physisporinus subcrocatus F. Wu, Jia J. Chen & Y.C. Dai, 新疆
621.Physisporinus tibeticus F. Wu, Jia J. Chen & Y.C. Dai, 西藏
622.Phylloporus brunneiceps N.K. Zeng & Zhu L. Yang, 云南
623.Phylloporus imbricatus N.K. Zeng & Zhu L. Yang, 云南
624.Phylloporus maculatus N.K. Zeng & Zhu L. Yang, 云南
625.Phylloporus pachycystidiatus N.K. Zeng & Zhu L. Yang, 云南
626.Phylloporus rubeolus N.K. Zeng & Zhu L. Yang, 云南
627.Phylloporus rubrosquamosus N.K. Zeng & Zhu L. Yang, 云南
628.Phylloporus yunnanensis N.K. Zeng & Zhu L. Yang, 云南
629.Physalacria corneri Zhu L. Yang & J. Qin, 云南
630.Physalacria lacrymispora Zhu L. Yang & J. Qin, 云南
631.Physalacria sinensis Zhu L. Yang & J. Qin, 云南
632.Picipes baishanzuensis J.L. Zhou & B.K. Cui, 浙江
633.Picipes hainanensis J.L. Zhou & B.K. Cui, 海南
634.Picipes jiajinensis J.L. Zhou & B.K. Cui, 四川
635.Picipes pseudovarius J.L. Zhou & B.K. Cui, 云南
636.Picipes subtropicus J.L. Zhou & B.K. Cui, 浙江
637.Picipes subtubaeformis J.L. Zhou & B.K. Cui, 安徽
638.Picipes tibeticus J.L. Zhou & B.K. Cui, 西藏
639.Piptoporellus hainanensis M.L. Han, B.K. Cui & Y.C. Dai, 海南
640.Piptoporellus triqueter M.L. Han, B.K. Cui, & Y.C. Dai, 云南
641.Podoserpula ailaoshanensis J.L. Zhou & B.K. Cui, 云南
642.Polyporus conifericola H.J. Xue & L.W. Zhou, 内蒙古
643.Polyporus cuticulatus Y.C. Dai, Jing Si & Schigel, 云南
644.Polyporus hapalopus H.J. Xue & L.W. Zhou, 广西
645.Polyporus mangshanensis B.K. Cui, J.L. Zhou & Y.C. Dai, 湖南
646.Polyporus pumilus Y.C. Dai & Niemelä, 湖南
647.Polyporus rhododendri Y.C. Dai & H.S. Yuan, 陕西
648.Polyporus submelanopus H.J. Xue & L.W. Zhou, 青海
649.Polyporus subvarius C.J. Yu & Y.C. Dai, 西藏
650.Polyporus taibaiensis Y.C. Dai, 陕西
651.Porodaedalea alpicola S.J. Dai, F. Wu & Y.C. Dai, 西藏
652.Porodaedalea chinensis S.J. Dai & F. Wu, 云南
653.Porodaedalea microsperma S.J. Dai, F. Wu & Y.C. Dai, 黑龙江
654.Porodaedalea mongolica Y.D. Wu & Y. Yuan 内蒙古
655.Porodaedalea yunnanensis S.J. Dai, F. Wu & Y.C. Dai, 云南
656.Porphyrellus castaneus Yan C. Li & Zhu L. Yang, 云南
657.Porphyrellus orientifumosipes Yan C. Li & Zhu L. Yang, 福建
658.Porpomyces submucidus F. Wu & C.L. Zhao, 海南
659.Postia amurensis Y.C. Dai & Penttila, 黑龙江
660.Postia amylocystis Y.C. Dai & Renvall, 吉林
661.Postia calcarea Y.L. Wei & Y.C. Dai, 安徽
662.Postia cana H.S. Yuan & Y.C. Dai, 山西
663.Postia cylindrica H.S. Yuan, 江西
664.Postia duplicata L.L. Shen, B.K. Cui & Y.C. Dai, 浙江
665.Postia gloeocystidiata Y.L. Wei & Y.C. Dai, 浙江
666.Postia gloeopora L.L. Shen, B.K. Cui & Y.C. Dai, 西藏
667.Postia hirsuta L.L. Shen & B.K. Cui, 陕西
668.Postia obliqua Y.L. Wei & W.M. Qin, 西藏
669.Postia ochraceoalba L.L. Shen, B.K. Cui & Y.C. Dai, 四川
670.Postia qinensis Y.C. Dai & Y.L. Wei, 陕西
671.Postia sublowei B.K. Cui, L.L. Shen & Y.C. Dai, 西藏
672.Postia subplacenta B.K. Cui, 吉林
673.Postia subundosa Y.L. Wei & Y.C. Dai, 黑龙江
674.Postia zebra Y.L. Wei & W.M. Qin, 吉林
675.Protomerulius efibulatus Y.C. Dai & Y.L. Wei, 海南
676.Pseudoarmillariella bacillaris Zhu L. Yang, B. Feng & Y.J. Hao, 四川
677.Pseudochaete latesetosa S.H. He & Hai J. Li, 海南
678.Pseudochaete subrigidula S.H. He & Hai J. Li, 云南
679.Pseudofibroporia citrinella Yuan Y. Chen & B.K. Cui, 广西
680.Pseudowrightoporia crassihypha Y.C. Dai, Jia J. Chen & B.K. Cui, 广西
681.Pseudowrightoporia hamata Y.C. Dai, Jia J. Chen & B.K. Cui, 湖南
682.Pseudowrightoporia oblongispora Y.C. Dai, Jia J. Chen & B.K. Cui, 广西
683.Psilocybe taiwanensis Zhu L. Yang & Guzmán, 台湾
684.Pulveroboletus brunneopunctatus G. Wu & Zhu L. Yang, 云南
685.Pulveroboletus flaviscabrosus N.K. Zeng & Zhu L. Yang, 云南
686.Pulveroboletus macrosporus G. Wu & Zhu L. Yang, 云南
687.Pulveroboletus rubroscabrosus N.K. Zeng & Zhu L. Yang, 湖北
688.Pulveroboletus subrufus N.K. Zeng & Zhu L. Yang, 海南
689.Pyrofomes castanopsidis B.K. Cui & Y.C. Dai, 广东
690.Pyrrhoderma hainanense L.W. Zhou & Y.C. Dai, 海南
691.Pyrrhoderma yunnanense L.W. Zhou & Y.C. Dai, 云南
692.Radulotubus resupinatus Y.C. Dai, S.H. He & C.L. Zhao, 云南
693.Ramaria luteoaeruginea P. Zhang, Zhu L. Yang & Ge, 青海
694.Ramaria pallidolilacina P. Zhang, Zhu L. Yang & Ge, 青海
695.Retiboletus brunneolus Yan C. Li & Zhu L. Yang, 福建
696.Retiboletus pseudogriseus N.K. Zeng & Zhu L. Yang, 福建
697.Retiboletus sinensis N.K. Zeng & Zhu L. Yang, 福建
698.Retiboletus zhangfeii N.K. Zeng & Zhu L. Yang, 福建
699.Rhodonia tianshanensis Yuan Yuan & L.L. Shen, 新疆
700.Rhodotus asperior L.P. Tang, Zhu L. Yang & B. Tolgor, 云南
701.Rigidoporopsis tegularis Juan Li & Y.C. Dai, 湖北
702.Rigidoporus eminens Y.C. Dai, 吉林
703.Rigidoporus fibulatus H.S. Yuan & Y.C. Dai, 广西
704.Rigidoporus minutus B.K. Cui & Y.C. Dai, 福建
705.Rigidoporus submicroporus F. Wu, Jia J. Chen & Y.C. Dai, 西藏
706.Rigidotubus tephroleucus J. Song, Y.C. Dai & B.K. Cui, 海南
707.Royoungia grisea Yan C. Li & Zhu L. Yang, 河南
708.Royoungia reticulata Yan C. Li & Zhu L. Yang, 云南
709.Royoungia rubina Yan C. Li & Zhu L. Yang, 福建
710.Rubroboletus latisporus Kuan Zhao & Zhu L. Yang, 重庆
711.Rugiboletus brunneiporus G. Wu & Zhu L. Yang, 西藏
712.Russula griseocarnosa X.H. Wang, Zhu L. Yang & Knudsen, 云南
713.Samsoniella alpina H. Yu, Y.B. Wang, Y. Wang & Zhu L. Yang, 云南
714.Samsoniella antleroides H. Yu, Y.B. Wang, Y. Wang, Q. Fan & Zhu L. Yang, 云南
715.Samsoniella cardinalis H. Yu, Y.B. Wang, Y. Wang, Q. Fan & Zhu L. Yang, 云南
716.Samsoniella cristata H. Yu, Y.B. Wang, Y. Wang, Q. Fan & Zhu L. Yang, 云南
717.Samsoniella kunmingensis H. Yu, Y.B. Wang, Y. Wang, Q. Fan & Zhu L. Yang, 云南
718.Samsoniella lanmaoa H. Yu, Y.B. Wang, Y. Wang, Q. Fan & Zhu L. Yang, 云南
719.Samsoniella ramosa H. Yu, Y.B. Wang, Y. Wang, Q. Fan & Zhu L. Yang, 云南
720.Samsoniella tortricidae H. Yu, Y.B. Wang, Y. Wang, Q. Fan & Zhu L. Yang, 云南
721.Samsoniella yunnanensis H. Yu, Y.B. Wang, Y. Wang, D.E. Duan & Zhu L. Yang, 云南
722.Sanghuangporus quercicola L. Zhu & B.K. Cui, 河南
723.Sanghuangporus toxicodendri Sheng H. Wu, B.K. Cui & Guo Z. Jiang, 湖北
724.Sanguinoderma laceratum Y.F. Sun & B.K. Cui, 云南
725.Sanguinoderma microporum Y.F. Sun & B.K. Cui, 海南
726.Sarcodon coactus Y.H. Mu & H.S. Yuan, 云南
727.Sarcodon grosselepidotus Y.H. Mu & H.S. Yuan, 云南
728.Sarcodon lidongensis Y.H. Mu & H.S. Yuan, 云南
729.Scutellinia pseudovitreola W.Y. Zhuang & Zhu L. Yang, 云南
730.Sidera parallela Y.C. Dai, F. Wu, G.M. Gates & Rui Du, 云南
731.Simplicillium yunnanense H. Yu, Y.B. Wang, Y. Wang & Zhu L. Yang, 云南
732.Singerocybe umbilicata Zhu L. Yang & J. Qin, 云南
733.Sinoboletus wangii M. Zang, Zhu L. Yang et Y. Zhang, 云南
734.Sistotrema subconfluens L.W. Zhou, 吉林
735.Skeletocutis bambusicola L.W. Zhou & W.M. Qin, 云南
736.Skeletocutis fimbriata Juan Li & Y.C. Dai, 湖北
737.Skeletocutis inflata B. K. Cui, 广东
738.Skeletocutis luteolus B.K. Cui & Y.C. Dai, 海南
739.Skeletocutis pseudo-odora L.F. Fan & Jing Si, 贵州
740.Skeletocutis substellae Y.C. Dai, 海南
741.Skeletocutis subvulgaris Y.C. Dai, 吉林
742.Skeletocutis yunnanensis L.S. Bian, C.L. Zhao & F. Wu, 云南
743.Sparassis cystidiosa f. flabelliformis Q. Zhao, Zhu L. Yang & Y.C. Dai, 云南
744.Sparassis latifolia Y.C. Dai & Zheng Wang, 吉林
745.Sparassis subalpina Q. Zhao, Zhu L. Yang & Y.C. Dai, 云南
746.Steccherinum cremicolor H.S. Yuan & Sheng H. Wu, 台湾
747.Steccherinum elongatus H.S. Yuan & Sheng H. Wu, 台湾
748.Steccherinum mukhinii Kotir. & Y.C. Dai, 吉林
749.Steccherinum subglobosum H.S. Yuan & Y.C. Dai, 湖北
750.Steccherinum subulatum H.S. Yuan & Y.C. Dai, 湖北
751.Stereum lithocarpi Y.C. Dai, 云南
752.Strobilomyces albidus L.H. Han, J. Xu & Zhu L. Yang, 云南
753.Strobilomyces anthracinus L.H. Han, J. Xu & Zhu L. Yang, 云南
754.Strobilomyces calidus L.H. Han, J. Xu & Zhu L. Yang, 云南
755.Strobilomyces cingulatus L.H. Han & Zhu L. Yang, 云南
756.Strobilomyces densisquamosus L.H. Han & Zhu L. Yang, 云南
757.Strobilomyces douformis L.H. Han & Zhu L. Yang, 云南
758.Strobilomyces microreticulatus L.H. Han & Zhu L. Yang, 云南
759.Strobilomyces pinophilus L.H. Han & Zhu L. Yang, 安徽
760.Strobilurus orientalis Zhu L. Yang & J. Qin, 云南
761.Strobilurus pachycystidiatus Zhu L. Yang & J. Qin, 云南
762.Subulicystidium acerosum S.H. He & S.L. Liu, 贵州
763.Subulicystidium tropicum S.H. He & S.L. Liu, 海南
764.Suillellus subamygdalinus Kuan Zhao & Zhu L. Yang, 西藏
765.Sutorius ferrugineus G. Wu, F. Li & Zhu L. Yang, 广东
766.Sutorius flavidus G. Wu & Zhu L. Yang, 云南
767.Sutorius rubriporus G. Wu & Zhu L. Yang, 云南
768.Sutorius sanguineoides G. Wu & Zhu L. Yang, 云南
769.Sutorius sanguineus G. Wu & Zhu L. Yang, 云南
770.Tengioboletus glutinosus G. Wu & Zhu L. Yang, 湖南
771.Tengioboletus reticulatus G. Wu & Zhu L. Yang, 湖南
772.Theleporus membranaceus Y.C. Dai & L.W. Zhou, 海南
773.Theleporus minisporus Y.C. Dai & L.W. Zhou, 海南
774.Theleporus rimosus H.S. Yuan, 广西
775.Tinctoporellus bubalinus H.S. Yuan, 广西
776.Tinctoporellus hinnuleus H.S. Yuan, 广西
777.Tomentella asiae-orientalis H.S. Yuan, X. Lu & Y.C. Dai, 黑龙江
778.Tomentella atrobadia H.S. Yuan & Y.C. Dai, 辽宁
779.Tomentella atrocastanea H.S. Yuan, X. Lu & Y.C. Dai, 辽宁
780.Tomentella aureomarginata H.S. Yuan, X. Lu & Y.C. Dai, 辽宁
781.Tomentella brevis H.S. Yuan, X. Lu & Y.C. Dai, 吉林
782.Tomentella brunneoflava H.S. Yuan, X. Lu & Y.C. Dai, 辽宁
783.Tomentella brunneogrisea H.S. Yuan, X. Lu & Y.C. Dai, 辽宁
784.Tomentella capitatocystidiata H.S. Yuan, X. Lu & Y.C. Dai, 吉林
785.Tomentella changbaiensis H.S. Yuan, X. Lu & Y.C. Dai, 吉林
786.Tomentella citrinocystidiata H.S. Yuan & Y.C. Dai, 辽宁
787.Tomentella coffeae H.S. Yuan & Y.C. Dai, 辽宁
788.Tomentella conclusa H.S. Yuan, X. Lu & Y.C. Dai, 黑龙江
789.Tomentella cystidiata H.S. Yuan & Y.C. Dai, 辽宁
790.Tomentella dimidiata H.S. Yuan, X. Lu & Y.C. Dai, 辽宁
791.Tomentella duplexa H.S. Yuan, X. Lu & Y.C. Dai, 辽宁
792.Tomentella efibulata H.S. Yuan & Y.C. Dai, 辽宁
793.Tomentella efibulis H.S. Yuan, X. Lu & Y.C. Dai, 辽宁
794.Tomentella farinosa H.S. Yuan & Y.C. Dai, 辽宁
795.Tomentella flavidobadia H.S. Yuan & Y.C. Dai, 辽宁
796.Tomentella fuscocrustosa H.S. Yuan, X. Lu & Y.C. Dai, 吉林
797.Tomentella fuscofarinosa H.S. Yuan, X. Lu & Y.C. Dai, 辽宁
798.Tomentella fuscogranulosa H.S. Yuan & Y.C. Dai, 辽宁
799.Tomentella fuscopelliculosa H.S. Yuan, X. Lu & Y.C. Dai, 吉林
800.Tomentella globospora H.S. Yuan & Y.C. Dai, 辽宁
801.Tomentella gloeocystidiata H.S. Yuan & Y.C. Dai, 辽宁
802.Tomentella griseocastanea H.S. Yuan, X. Lu & Y.C. Dai, 吉林
803.Tomentella griseofusca H.S. Yuan & Y.C. Dai, 辽宁
804.Tomentella griseomarginata H.S. Yuan, X. Lu & Y.C. Dai, 吉林
805.Tomentella inconspicua H.S. Yuan & Y.C. Dai, 辽宁
806.Tomentella incrustata H.S. Yuan, X. Lu & Y.C. Dai, 辽宁
807.Tomentella interrupta H.S. Yuan & Y.C. Dai, 辽宁
808.Tomentella liaoningensis H.S. Yuan & Y.C. Dai, 辽宁
809.Tomentella longiaculeifera H.S. Yuan & Y.C. Dai, 辽宁
810.Tomentella longiechinuli H.S. Yuan, X. Lu & Y.C. Dai, 黑龙江
811.Tomentella megaspora H.S. Yuan, X. Lu & Y.C. Dai, 吉林
812.Tomentella olivacea H.S. Yuan & Y.C. Dai, 辽宁
813.Tomentella olivaceobrunnea H.S. Yuan, X. Lu & Y.C. Dai, 辽宁
814.Tomentella pallidobrunnea H.S. Yuan, X. Lu & Y.C. Dai, 吉林
815.Tomentella pallidocastanea X. Lu, Y.H. Mu & H.S. Yuan, 黑龙江
816.Tomentella pallidomarginata H.S. Yuan, X. Lu & Y.C. Dai, 吉林
817.Tomentella parvispora H.S. Yuan & Y.C. Dai, 辽宁
818.Tomentella pertenuis H.S. Yuan & Y.C. Dai, 辽宁
819.Tomentella qingyuanensis H.S. Yuan & Y.C. Dai, 辽宁
820.Tomentella segregata H.S. Yuan, X. Lu & Y.C. Dai, 辽宁
821.Tomentella separata H.S. Yuan, X. Lu & Y.C. Dai, 辽宁
822.Tomentella stipitata H.S. Yuan, X. Lu & Y.C. Dai, 辽宁
823.Tomentella storea H.S. Yuan & Y.C. Dai, 辽宁
824.Tomentella tenuirhizomorpha X. Lu, Y.H. Mu & H.S. Yuan, 黑龙江
825.Trametes cystidiolophora B.K. Cui & H.J. Li, 云南
826.Trametes duplexa Hai J. Li, Y.C. Dai & B.K. Cui, 海南
827.Trametes ellipsoidea Hai J. Li, Y.C. Dai & B.K. Cui, 云南
828.Trametes stipitata Hai J. Li, Y.C. Dai & B.K. Cui, 云南
829.Trametes subsuaveolens B.K. Cui & Y.C. Dai, 内蒙古
830.Trechispora suberosa H.S. Yuan & Y.C. Dai, 广西
831.Trichaptum imbricatum Y.C. Dai & B.K. Cui, 海南
832.Trichaptum perenne Y.C. Dai & H.S. Yuan, 云南
833.Trichaptum podocarpi Y.C. Dai, 海南
834.Tricholoma forteflavescens K. Reschke, F. Popa, Z.L. Yang & G. Kost, 云南
835.Tricholoma highlandense Zhu L Yang, X.X. Ding, G. Kost & Rexer, 云南
836.Tricholoma melleum K. Reschke, F. Popa, Z.L. Yang & G. Kost, 云南
837.Tricholoma muscarioides K. Reschke, F. Popa, Zhu L. Yang & G. Kost, 云南
838.Tricholoma olivaceoluteolum K. Reschke, F. Popa, Zhu L. Yang & G. Kost, 云南
839.Tricholoma olivaceum K. Reschke, F. Popa, Z.L. Yang & G. Kost, 云南
840.Tricholoma orienticolossus X. Xu, Y.Y. Cui & Zhu L. Yang, 四川
841.Tricholoma orientifulvum X. Xu, Y.Y. Cui & Zhu L. Yang, 云南
842.Tricholoma sinopardinum Zhu L Yang, X.X. Ding, G. Kost & Rexer, 西藏
843.Tricholoma sinoportentosum Zhu L. Yang, K. Reschke, F. Popa & G. Kost, 云南
844.Tricholyophyllum brunneum Qing Cai, G. Kost & Zhu L. Yang, 云南
845.Trogia venenata Zhu L. Yang, Y.C. Li & L.P. Tang, 云南
846.Tropicoporus excentrodendri L.W. Zhou & Y.C. Dai, 广西
847.Truncospora macrospora B.K. Cui & C.L. Zhao, 云南
848.Tylocinum griseolum Yan C. Li & Zhu L. Yang, 云南
849.Tylopilus alpinus Yan C. Li & Zhu L. Yang, 云南
850.Tylopilus atroviolaceobrunneus Yan C. Li & Zhu L. Yang, 云南
851.Tylopilus violaceobrunneus Yan C. Li & Zhu L. Yang, 山东
852.Tyromyces minutulus Y.C. Dai & C.L. Zhao, 海南
853.Vararia bispora S.L. Liu & S.H. He, 广西
854.Vararia montana S.L. Liu & S.H. He, 云南
855.Veloporphyrellus alpinus Yan C. Li & Zhu L. Yang, 云南
856.Veloporphyrellus gracilioides Yan C. Li & Zhu L. Yang, 四川
857.Veloporphyrellus pseudovelatus Yan C. Li & Zhu L. Yang, 云南
858.Veluticeps fasciculata Jiao Yang, Nakasone & S.H. He, 江西
859.Veluticeps microspora S.H. He & Hai J. Li, 云南
860.Veluticeps nakasoneae Jiao Yang & S.H. He, 云南
861.Wolfiporia castanopsidis Y.C. Dai, 云南
862.Wolfiporia curvispora Y.C. Dai, 吉林
863.Wolfiporia pseudococos F. Wu, J. Song & Y.C. Dai, 海南
864.Wrightoporia austrosinensis Y.C. Dai, 海南
865.Wrightoporia biennis Jia J. Chen & B.K. Cui, 云南
866.Wrightoporia casuarinicola Y.C. Dai & B.K. Cui, 广西
867.Wrightoporia luteola B.K. Cui & Y.C. Dai, 安徽
868.Wrightoporia rubella Y.C. Dai, 北京
869.Wrightoporia subavellanea Jia J. Chen & B.K. Cui, 广西
870.Wrightoporia unguliformis Y.C. Dai & B.K. Cui, 海南
871.Wrightoporiopsis amylohypha Y.C. Dai, Jia J. Chen & B.K. Cui, 云南
872.Wrightoporiopsis irregularis YC Dai, Q Chen & XH Ji, 海南
873.Xanthoconium sinense G. Wu, Yang-Yang Cui & Zhu L. Yang, 广东
874.Xerocomellus communis Xue T. Zhu & Zhu L. Yang, 云南
875.Xerocomus fraternus Xue T. Zhu & Zhu L. Yang, 云南
876.Xerocomus fulvipes Xue T. Zhu & Zhu L. Yang, 云南
877.Xerocomus subparvus Xue T. Zhu & Zhu L. Yang, 云南
878.Xerocomus velutinus Xue T. Zhu & Zhu L. Yang, 云南
879.Xerula strigosa Zhu L. Yang, L. Wang & G.M. Muller, 云南
880.Xylobolus austrosinensis S.H. He, 江西
881.Xylodon kunmingensis L.W. Zhou & C.L. Zhao, 云南
882.Zangia citrina Yan C. Li & Zhu L. Yang, 福建
883.Zangia erythrocephala Yan C. Li & Zhu L. Yang, 云南
884.Zangia olivacea Yan C. Li & Zhu L. Yang, 云南
885.Zangia olivaceobrunnea Yan C. Li & Zhu L. Yang, 云南
3 新种涉及高等真菌各大主要纲目
研究发现的885个新种隶属于真菌界的2个门(子囊菌门和担子菌门)、5个纲(盘菌纲Pezizomycetes、粪壳菌纲Sordariomycetes、伞菌纲Agaricomycetes、花耳纲Dacrymycetes和银耳纲Tremellomycetes)、17个目、63个科、247个属,另有5个新种的科级分类位置不定。其中担子菌门种类861种,占97.3%;伞菌纲种类有859种,占97%,因此绝大多数新种为担子菌门伞菌纲的种类。885个新种在目级分类单元的分布见图1,其中多孔菌目、锈革孔菌目、伞菌目、牛肝菌目、革菌目和红菇目分别有280、187、156、99、55和54种,分别占新种总数的31.6%、21.1%、17.6%、11.2%、6.2%和6.1%,这6个目包括的新种占全部新种的94%。在科级水平上锈革孔菌科、多孔菌科、牛肝菌科、鹅膏菌科Amanitaceae、革菌科Thelephoraceae和齿耳科Steccherinaceae分别有163、131、95、84、52和31种,分别占新种总数的18%、11%、10%、9%、6%和4%,这6个科包括的新种占全部新种的63%。上述数据表明在这些目和科中我国的大型真菌多样性较为丰富,研究也较为深入。
图1
图1
森林大型真菌重要类群885个新种在目级分类单元的分布
Fig. 1
The distribution at order level of 885 new macrofungi from Chinese forests.
4 新种主模式产地覆盖全国30个省区
图2
图2
大型真菌重要类群885个新种主模式产地的省级分布
Fig. 2
The provincial distribution of the holotypes of 885 new macrofungi from Chinese forests.
5 我国重要食药用真菌名称得到及时更新
我国最著名的药用真菌灵芝从1907年(Patouillard 1907)被鉴定为欧洲灵芝Ganoderma lucidum (Curtis) P. Karst.后,一直到2012年才被证实它与欧洲灵芝不同,为一个新种,并以灵芝的汉语拼音命名为G. lingzhi(Cao et al. 2012)。无独有偶,在19世纪后期Kalchbrenner & Thümen(1881)将我国著名的食用菌黑木耳鉴定为欧洲木耳 Auricularia auricula (L.) Underw = A. auricula-judae (Bull.) Quél.,一直到2014年才被证实中国的黑木耳与欧洲木耳是两个不同的物种,故黑木耳被命名为A. heimuer (Wu et al. 2014)。研究还发现,在东亚地区广泛栽培的冬菇(俗称“金针菇”)过去一直认为是欧洲毛腿冬菇Flammulina velutipes (Curtis) Singer,但实际上东亚的冬菇和欧洲的毛腿冬菇是不同的物种,最近才被更新为F. filiformis (Z.W. Ge, X.B. Liu & Zhu L. Yang) P.M. Wang, Y.C. Dai, E. Horak & Zhu L. Yang(戴玉成和杨祝良2018;Wang et al. 2018)。此外,对我国药用真菌物种多样性进行全面更新,中国药用真菌有540种,最主要的药用功能为增强免疫力、抗肿瘤、抗氧化和抗细菌(戴玉成和杨祝良 2008;Dai et al. 2009)。
6 毒蘑菇研究成果支撑全国毒蘑菇中毒防治工作
2000年3月,9名务工人员在广州误食野生白色蘑菇,导致8人中毒死亡,经过研究发现该毒蘑菇是个新种,并命名为致命鹅膏Amanita exitialis(Yang & Li 2001)。致命鹅膏在我国至少引发了30余起中毒事件,造成100余人中毒,其中超过60人死亡,是我国最著名的剧毒蘑菇之一(陈作红等 2016)。此外,研究还发现了拟灰花纹鹅膏Amanita fuligineoides(Zhang et al. 2010)、灰盖粉褶鹅膏Amanita griseorosea(Cai et al. 2016)、浅红鹅膏Amanita pallidorosea(Zhang et al. 2010)、裂皮鹅膏Amanita rimosa(Zhang et al. 2010)、毒环柄菇Lepiota venenata、毒沟褶菌Trogia venenata等剧毒蘑菇。在我国的大型真菌中,共有毒蘑菇480种,对子实体发生时间和地理分布特征有了较为深入的认识(Wu et al. 2019),为毒蘑菇准确鉴定提供了重要依据,对预防和治疗蘑菇中毒具有重要现实意义。
7 我国森林病原真菌物种多样性得到系统论述
研究过程中共发现造成我国木本植物腐朽病害的病原菌152种(戴玉成 2012),其中栎生多孢孔菌Abundisporus quercicola Y.C. Dai和罗汉松邦氏孔菌Bondarzewia podocarpi Y.C. Dai & B.K. Cui等49种(占总数的34%)为研究过程中在我国首次报道为林木病原菌。对每个病原菌的寄主、侵染部位、腐朽类型、发生频率及分布进行了报道。135种(占总数的89%)造成木材白色腐朽,17种(占总数的11%)造成褐色腐朽。对世界范围内最严重森林病害即针叶树干基腐朽病的病原菌进行了系统研究,在我国针叶树干基腐朽病的病原菌过去普遍被认为是多年异担子菌Heterobasidion annosum (Fr.) Bref.,该真菌最初发现于欧洲,过去基于传统的分类鉴定方法认为该病原菌在我国东北和西南等地区以及国外的美洲地区也广泛分布(袁嗣令 1997)。最新的研究发现我国广泛分布的异担子菌种类实际上是亚小孔异担子菌H. subparviporum,而真正的多年异担子菌在我国并没有分布(Yuan et al. 2021)。世界范围内能够引起针叶树干基腐朽病的病原菌包括欧洲的冷杉异担子菌H. abietinum Niemelä & Korhonen、小孔异担子菌H. parviporum Niemelä & Korhonen和多年异担子菌及美洲的西方异担子菌H. occidentale Otrosina & Garbel.和变孔异担子菌H. irregulare Garbel. & Otrosina,这几种造成针叶树干基腐朽病严重危害的种类在我国并没有分布,应将这些种类列为重要入境检疫对象(Yuan et al. 2021)。
8 森林大型真菌代表性类群的起源和演化规律始有认识
异担子菌属Heterobasidion种类引起世界范围内针叶树最严重的森林病害。基于多基因片段数据,运用最大似然法和贝叶斯法对该属的12个种进行了系统发育关系分析,发现异担子菌属可能起源于中新世早期的欧亚大陆,随后在中新世中期和上新世早期扩散至其他大陆并形成物种。变孔异担子菌和西方异担子菌分别越过白令海峡和北大西洋路桥在北美成种,南半球南洋杉异担子菌H. araucariae P.K. Buchanan可能是由于最近人类迁徙引入的。板块构造和长距离扩散是影响异担子菌成种和生物地理分布的最可能因素(Chen et al. 2015)。
檐托鹅膏组Amanita sect. Phalloideae (Fr.) Quél.是鹅膏属中较为独特的一个类群,因其含有致命的环肽类毒素而闻名。综合形态学、解剖学和生态学特征及相关生物化学分析研究,作者构建了全球剧毒鹅膏较为全面的分子系统发育框架,共界定了剧毒鹅膏28个系统发育种,研究表明剧这些剧毒鹅膏是一个自然的单系类群,鹅膏毒肽(amatoxins)和鬼笔毒肽(phallotoxins)为单次起源,是剧毒鹅膏的一个共衍征。研究显示剧毒鹅膏起源于古新世早期的古热带地区,在渐新世中期至中新世期间通过白令陆桥传播至北美,后由于中新世后期全球气候变冷,白令陆桥的阻断而发生异域成种事件。扩散和隔离分化在剧毒鹅膏的洲际间断分布格局形成过程中起了重要作用,东亚为剧毒鹅膏现代分布中心和物种多样性中心(Cai et al. 2014)。
松塔牛肝菌属Strobilomyces Berk.真菌为世界广布的外生菌根菌,在全球广泛取样的基础上,通过多基因片段构建了该属较为全面的分子系统发育框架。基于分子钟推算、祖先分布区和宿主植物重建,研究了该属的生物地理,发现该属约在始新世起源于非洲,之后借助“北方热带森林”扩散至东南亚,形成了非洲-东南亚古热带间断分布格局。在渐新世期间从东南亚向北别扩散至东亚、向南扩散至大洋洲。中新世期间通过白令陆桥和中国东海陆桥迁移以及气候变化驱动的隔离分化等在北半球形成间断分布格局。在中新世中期,东亚发生了一次快速辐射演化事件,致使其成为该属的现代分布中心和物种多样性中心(Han et al. 2017)。
糙皮侧耳复合群Pleurotus ostreatus (Jacq.) P. Kumm. complex是具有重要经济和生态意义的腐生真菌。基于全球样品和40个核单拷贝同源基因,运用最大似然法和贝叶斯法对该类群进行系统发育关系分析,共界定了糙皮侧耳复合群20个系统发育种。分子钟分析数据表明,侧耳属的分化可能发生在侏罗纪晚期,而糙皮侧耳复合群的共同祖先可能在始新世末(约39Mya)出现于东亚。在晚渐新世、晚中新世和晚上新世期间糙皮侧耳的种类可能通过北大西洋大陆桥或白令海峡大陆桥分别从东亚迁移至北美,然后通过多次分散和分隔事件在新旧世界同时分化。晚中新世和第四纪冰期循环,干旱加剧和气候变冷事件对北半球物种多样性分化产生了深刻影响(Li et al. 2020)。
9 初步揭示了我国森林大型真菌分布格局
深入分析和系统总结前人研究我国大型真菌及其他地区真菌重要论著成果后,发现我国现在大型真菌区系具有复杂而独特的性质,提出了我国大型真菌呈现出“东喜马拉雅-横断山区”、“华中地区”和“中国热带地区”三大物种多样性中心的分布格局,论证了该格局应是地质历史事件、地理因素、多样化地理成分、新老种汇聚、与植物协同进化和复杂的生态环境等多种因子共同作用的结果。复杂性主要表现为其与世界其他区系如非洲、北美洲、大洋洲有各种联系,融入了多种地理起源迥异的成分;该区系中还汇聚了古老种、孑遗种和新生种等成分。独特性主要表现为该区系3个中心分别具有各自独特的真菌属和种(Yang 2005)。
10 研究展望
研究人员虽然取得了上述研究成果,但尚有许多类群还有待进一步研究。一方面,除了森林生态系统外,还有草原生态系统、荒漠生态系统及农田生态系统的大型真菌资源和多样性研究需要进一步展开。另一方面,今后研究中还需要聚焦在以下几个重点领域:(1)加强薄弱类群和区域大型真菌的深入和系统研究;(2)加强大型真菌DNA条形码研究,以便非专业人士快速鉴别大型真菌物种;(3)对重要类群开展国际合作研究,强强联合,不断提高研究水平;(4)从系统发育基因组角度,研究重要类群的科、目甚至纲级的系统进化关系。
深入认识我国乃至东亚和东南亚的大型真菌多样性的起源和演化规律,可为资源开发和利用奠定科学基础,为揭示森林生态系统物种多样性维持机制提供科学支撑。
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