高寒草甸不同放牧强度下植物地上-地下生物量权衡的变化

doi:10.11733/j.issn.1007-0435.2025.08.018

草地学报 ›› 2025, Vol. 33 ›› Issue (8): 2575-2584.DOI: 10.11733/j.issn.1007-0435.2025.08.018

高寒草甸不同放牧强度下植物地上-地下生物量权衡的变化

段俊光¹, 房凯¹, 裴渌², 褚建民¹, 张金鑫², 李晓霞², 王迎新²

1. 中国林业科学研究院林业研究所, 北京 100091;
2. 中国林业科学研究院生态保护与修复研究所/国家林业和草原局草原研究中心, 北京 100091

收稿日期:2025-04-02 修回日期:2025-05-19 出版日期:2025-08-15 发布日期:2025-08-27
通讯作者: 王迎新, E-mail:wyxcaf@163.com
作者简介:#段俊光(2000-),男,汉族,湖南岳阳人,硕士研究生,主要从事退化草地修复研究,E-mail:DUANJG2000@163.com;房凯(1991-),男,汉族,山东德州人,博士,助理研究员,主要从事土壤生态学研究,E-mail:fangkai@caf.ac.cn
基金资助:
中央级公益性科研院所基本科研业务费专项资金（CAFYBB2024MA010， CAFYBB2020ZB001， CAFYBB2023ZA009-03， CAFYBB2022XA002）；中国科学院战略性先导科技专项（A类）（XDA26020102）资助

Changes of Trade-offs Between Aboveground and Belowground Plant Biomass Across Different Grazing Intensities of Alpine Meadow

DUAN Jun-guang¹, FANG Kai¹, PEI Lu², CHU Jian-min¹, ZHANG Jin-xin², LI Xiao-xia², WANG Ying-xin²

1. Research Institute of Forestry, Chinese Academy of Forestry, Beijing 100091, China;
2. Institute of Ecological Conservation and Restoration, Chinese Academy of Forestry, Beijing 100091, China

Received:2025-04-02 Revised:2025-05-19 Online:2025-08-15 Published:2025-08-27

摘要/Abstract

摘要： 近年来草地过度放牧现象日益加剧，对高寒草甸生态系统的稳定性造成严重威胁。明确不同放牧强度下植物地上-地下生物量的权衡关系及其驱动因素，对高寒草甸生态保护与恢复具有重要意义。本研究以青藏高原红原县无放牧、轻度、中度和重度放牧的高寒草甸为对象，通过测定植物地上与地下生物量及土壤理化性质，探究不同放牧强度下植物生物量分配的权衡变化及其驱动因素。结果显示：无放牧草地生物量向地下权衡，随放牧强度增加，生物量逐渐向地上倾斜，权衡值降低。生物量权衡与植物株高、物种丰富度、土壤全碳及全氮含量呈显著正相关关系，与土壤碳氮比、容重呈显著负相关关系。方差分解表明，轻度放牧阶段植物特性是主要驱动因素，中度和重度放牧阶段土壤因素的作用更明显。因此，在退化高寒草甸的管理实践中，应根据放牧强度的差异，针对性地调整植被管理策略和土壤养分恢复措施，以实现生态系统的有效恢复。

关键词: 高寒草甸, 放牧强度, 生物量权衡, 植物特性, 土壤养分

Abstract: In recent decades， the exacerbation of overgrazing has posed substantial threats to the stability of alpine meadow ecosystems. Understanding the trade-offs between aboveground and belowground biomass allocation under different grazing intensities， as well as identifying the driving factors behind these patterns， is critical for ecological conservation and restoration initiatives in alpine meadows. This study centered on alpine meadow ecosystems in Hongyuan County on the Tibetan Plateau， which were subjected to four distinct grazing intensity treatments： ungrazed， lightly grazed， moderately grazed， and heavily grazed. Through meticulous measurement of plant aboveground and belowground biomass， and analysis of soil physicochemical properties， we investigated the changes and driving factors behind biomass allocation under different grazing intensities. The results demonstrated that under ungrazed treatment， biomass allocation was skewed towards the belowground. However， as the grazing intensity increased， there was a progressive shift towards aboveground biomass， resulting in a decrease in the trade-off index. Plant height， plant species richness， soil total carbon， and soil total nitrogen content showed significant positive correlations with the biomass trade-off， whereas soil carbon-nitrogen ratio （C∶N） and soil bulk density were significantly and negatively correlated with the biomass trade-off. Variance partitioning analysis revealed that under light grazing conditions， plant characteristics were the primary drivers of biomass allocation trade-offs. Nevertheless， as the grazing intensity increased to moderate and heavy levels， soil factors became more influential. Consequently， management practices aimed at the restoration of degraded alpine meadows should be tailored specifically according to grazing intensity. This involves adjusting vegetation management strategies and implementing targeted measures to restore soil nutrients and effectively promote ecosystem restoration.

Key words: Alpine meadow, Grazing intensity, Biomass trade-off, Plant traits, Soil nutrients

中图分类号:

S812

段俊光, 房凯, 裴渌, 褚建民, 张金鑫, 李晓霞, 王迎新. 高寒草甸不同放牧强度下植物地上-地下生物量权衡的变化[J]. 草地学报, 2025, 33(8): 2575-2584.

DUAN Jun-guang, FANG Kai, PEI Lu, CHU Jian-min, ZHANG Jin-xin, LI Xiao-xia, WANG Ying-xin. Changes of Trade-offs Between Aboveground and Belowground Plant Biomass Across Different Grazing Intensities of Alpine Meadow[J]. Acta Agrestia Sinica, 2025, 33(8): 2575-2584.

参考文献

[1] BAR-ON Y M，PHILLIPS R，MILO R. The biomass distribution on earth[J]. Proceedings of the National Academy of Sciences of the United States of America，2018，115（25）：6506-6511
[2] POST A K，KNAPP A K. The importance of extreme rainfall events and their timing in a semi-arid grassland[J]. Journal of Ecology，2020，108（6）：2431-2443
[3] DING L，LI Z W，SHEN B B，et al. Spatial patterns and driving factors of aboveground and belowground biomass over the eastern Eurasian steppe[J]. Science of The Total Environment，2022，803：149700
[4] WANG X X，CHEN X H，XU J L，et al. Precipitation dominates the allocation strategy of above-and belowground biomass in plants on macro scales[J]. Plants，2023，12（15）：2843
[5] YANG L S，LI S Y，ZHENG J H，et al. Plant community biomass trade-off and its influencing factors for different years of abandoned cropland in a typical steppe[J]. Chinese Journal of Grassland，2024，46（4）：33-42 杨立山，李邵宇，郑佳华，等.典型草原区不同年限弃耕地植物群落生物量权衡及其影响因素[J].中国草地学报，2024，46（4）：33-42
[6] SUN J，MA B B，LU X Y. Grazing enhances soil nutrient effects： trade-offs between aboveground and belowground biomass in alpine grasslands of the Tibetan Plateau[J]. Land Degradation & Development，2018，29（2）：337-348
[7] DONG S K，ZHANG Y H，WANG G C. Assessment of grassland health and degradation： concepts， principles， and methods[J]. Pratacultural Science，2023，40（12）：2971-2981 董世魁，张宇豪，王冠聪.草地健康与退化评价：概念、原理及方法[J].草业科学，2023，40（12）：2971-2981
[8] ZHENG K，WEI J Z，PEI J Y，et al. Impacts of climate change and human activities on grassland vegetation variation in the Chinese Loess Plateau[J]. Science of the Total Environment，2019，660：236-244
[9] SHUKLA P R，SKEG J，CALVO BUENDIA E C，et al. Climate change and land： an IPCC special report on climate change， desertification， land degradation， sustainable land management， food security， and greenhouse gas fluxes in terrestrial ecosystems[J]. Intergovernmental Panel on Climate Change， 2019：45-84
[10] PARSONS P A. Environments and evolution： interactions between stress， resource inadequacy and energetic efficiency[J]. Biological Reviews，2005，80（4）：589-610
[11] YAN Y，ZHU J J，ZHANG B，et al. A review of belowground biomass allocation and its response to global climatic change in grassland ecosystems[J]. Chinese Journal of Plant Ecology，2017，41（5）：585-596 严月，朱建军，张彬，等.草原生态系统植物地下生物量分配及对全球变化的响应[J].植物生态学报，2017，41（5）：585-596
[12] MAN X Y，GUO Y R，GE H J，et al. Advances in mutual regulation mechanisms between below-and above-ground of plants[J]. Plant Physiology Journal，2018，54（6）：931-942 满欣雨，郭亚蓉，葛红娟，等.植物地下部与地上部远端相互调控研究进展[J].植物生理学报，2018，54（6）：931-942
[13] CUI B W，WANG X M，SU Y B，et al. Responses of biomass of urban trees to pavement and spacing[J]. Acta Ecologica Sinica，2022，42（11）：4463-4472 崔博文，汪旭明，苏跃波，等.城市林木生物量对硬化地表和种植密度的响应[J].生态学报，2022，42（11）：4463-4472
[14] BRADFORD J B，D'AMATO A W. Recognizing trade-offs in multi-objective land management[J]. Frontiers in Ecology and the Environment，2012，10（4）：210-216
[15] DONG Q M，ZHAO X Q，LIU Y Z，et al. Effects of different herbivore assemblage on relationship between Kobresia humilis seed size and seed number in an alpine grassland[J]. Chinese Journal of Plant Ecology，2022，46（9）：1018-1026 董全民，赵新全，刘玉祯，等.放牧方式影响高寒草地矮生嵩草种子大小与数量的关系[J].植物生态学报，2022，46（9）：1018-1026
[16] WU J H，LI Y，WANG Y X，et al. Trade-off between aboveground and belowground net primary productivity of plant communities in alpine grassland[J]. Acta Ecologica Sinica，2024，44（2）：793-804 吴江航，李洋，王迎新，等.高寒草地植物群落地上-地下净初级生产力权衡[J].生态学报，2024，44（2）：793-804
[17] GAO X X，DONG S K，XU Y D，et al. Plant biomass allocation and driving factors of grassland revegetation in a Qinghai-Tibetan Plateau chronosequence[J]. Land Degradation & Development，2021，32（4）：1732-1741
[18] LIU Y Z，ZHAO X Q，LIU W T，et al. Plant biomass partitioning in alpine meadows under different herbivores as influenced by soil bulk density and available nutrients[J]. Catena，2024，240：108017
[19] GAO X Y，LU X Y. Effects of rest grazing on vegetation and soil characteristics of alpine steppe and alpine meadow in Tibet[J]. Pratacultural Science，2020，37（3）：486-496 高小源，鲁旭阳.休牧对西藏高寒草原和高寒草甸植被与土壤特征的影响[J].草业科学，2020，37（3）：486-496
[20] LIU J Z，DONG R，HUA L M，et al. Effects of plateau zokor Eospalax baileyi disturbance on the aboveground-belowground biomass allocation pattern of plant communities and soil physicochemical properties in alpine meadow[J]. Journal of Plant Protection，2024，51（5）：1090-1098 刘济泽，董瑞，花立民，等.高原鼢鼠干扰对高寒草甸植物群落地上-地下生物量分配格局和土壤理化性质的影响[J].植物保护学报，2024，51（5）：1090-1098
[21] CHANG Y R，SHI J M，XU D M，et al. Trade-off relationships between biomass and nutrient allocation in different natural populations of Agropyron mongolicum on the desert steppe[J]. Acta Prataculturae Sinica，2024，33（11）：186-197 常怡然，史佳梅，许冬梅，等.荒漠草原不同自然种群蒙古冰草生物量和养分权衡特征[J].草业学报，2024，33（11）：186-197
[22] ZHAO W L，QI J G，SUN G J，et al. Spatial patterns of top soil carbon sensitivity to climate variables in northern Chinese grasslands[J]. Acta Agriculturae Scandinavica， Section B—Soil & Plant Science，2012，62（8）：720-731
[23] SUN J，ZHOU T C，LIU M，et al. Water and heat availability are drivers of the aboveground plant carbon accumulation rate in alpine grasslands on the Tibetan Plateau[J]. Global Ecology and Biogeography，2020，29（1）：50-64
[24] WANG X D，CHENG G W，ZHAO T，et al. Assessment on protection and construction of ecological safety shelter for Tibet[J]. Bulletin of Chinese Academy of Sciences，2017，32（1）：29-34 王小丹，程根伟，赵涛，等.西藏生态安全屏障保护与建设成效评估[J].中国科学院院刊，2017，32（1）：29-34
[25] WANG Y X，SUN Y，LIU Y，et al. Ecological thresholds of toxic plants for sheep production and ecosystem multifunctionality and their trade-off in an alpine meadow[J]. Journal of Environmental Management，2022，323：116167
[26] HUANG X J，HOU F J. Principle， technique and application of grassland improvement[J]. Journal of Environmental Management，2024，369：122264
[27] REN J Z，HU Z Z，ZHAO J，et al. A grassland classification system and its application in China[J]. The Rangeland Journal，2008，30（2）：199
[28] PEI L，WU Z，QIAN Y Q，et al. Plant community stability， indicator species and their driving factors at a gradient of grazing intensity in an alpine meadow[J]. Ecological Indicators，2024，162：112012
[29] LI H. Effects of long-term enclosure and grazing intensity on spatial heterogeneity of plant diversity and soil nutrient in typical steppe[D]. Hohhot：Inner Mongolia University，2021：10-11 李浩.长期围封与放牧对典型草原群落和土壤空间异质性的影响[D].呼和浩特：内蒙古大学，2021：10-11
[30] BARGAGLI R，BROWN D H，NELLI L. Metal biomonitoring with mosses： procedures for correcting for soil contamination[J]. Environmental Pollution，1995，89（2）：169-175
[31] ZHAGN X L，HU Z Q，CHU S L. Method for measuring soil water content： a review[J]. Chinese Journal of Soil Science，2005，（1）：118-123 张学礼，胡振琪，初士立.土壤含水量测定方法研究进展[J].土壤通报，2005，（1）：118-123
[32] ZHOU T C，SUN J，ZONG N，et al. Community species diversity mediates the trade‐off between aboveground and belowground biomass for grasses and forbs in degraded alpine meadow， Tibetan Plateau[J]. Ecology and Evolution，2021，11（19）：13259-13267
[33] PERES-NETO P R，LEGENDRE P，DRAY S，et al. Variation partitioning of species data matrices： estimation and comparison of fractions[J]. Ecology，2006，87（10）：2614-2625
[34] CHENG D L，NIKLAS K J. Above-and below-ground biomass relationships across 1534 forested communities[J]. Annals of Botany，2007，99（1）：95-102
[35] SUN L，SUN J，WU J X，et al. Plant community traits and functions mediate the biomass trade-off of alpine grasslands along precipitation gradients on the Tibetan Plateau[J]. Journal of Plant Ecology，2023，16（5）：rtad009
[36] XIA S M，WEI H R，ZHENG W，et al. Effects of different grazing methods on the production and ecological functions of mountain meadows and their trade-off relationships[J]. Acta Agrestia Sinica，2022，30（8）：1972-1981 夏树淼，蔚恒瑞，郑伟，等.不同放牧方式对山地草甸生产与生态功能及其权衡关系的影响[J].草地学报，2022，30（8）：1972-1981
[37] SUN J， WANG H M. Soil nitrogen and carbon determine the trade-off of the above-and below-ground biomass across alpine grasslands， Tibetan Plateau[J]. Ecological Indicators，2016，60：1070-1076
[38] ZHANG B，LI S Y，GU C，et al. Biomass allocation of four dominant plant species in Inner Mongolia desert grasslands in response to different grazing intensities [J]. Acta Agrestia Sinica， 2022，30（12）：3355-3363 张彬，李邵宇，古琛，等.内蒙古荒漠草原4种优势植物生物量分配对不同放牧强度的响应[J].草地学报，2022，30（12）：3355-3363
[39] ZENG C X，WU J S，ZHANG X Z. Effects of grazing on above- vs. below-ground biomass allocation of alpine grasslands on the northern Tibetan Plateau[J]. PLoS One，2015，10（8）：e0135173
[40] CHAPIN III F S，MATSON P A，VITOUSEK P. Principles of terrestrial ecosystem ecology[M]. 2th ed. Berlin： Springer Science & Business Media，2011：63-90
[41] TILMAN D. Plant strategies and the dynamics and structure of plant communities[M]. Princeton： Princeton University Press，1988：184-212
[42] LOREAU M，NAEEM S，INCHAUSTI P. Biodiversity and ecosystem functioning： synthesis and perspectives[M]. New York： Oxford University Press，2002：79-101
[43] HECTOR A，SCHMID B，BEIERKUHNLEIN C，et al. Plant diversity and productivity experiments in European grasslands[J]. Science，1999，286（5442）：1123-1127
[44] JOBBÁGY E G，JACKSON R B. The vertical distribution of soil organic carbon and its relation to climate and vegetation[J]. Ecological Applications，2000，10（2）：423-436
[45] SIX J，CONANT R T，PAUL E A， et al. Stabilization mechanisms of soil organic matter： implications for C-saturation of soils[J]. Plant and Soil，2002，241（2）：155-176
[46] WARDLE D A，BARDGETT R D，KLIRONOMOS J N，et al. Ecological linkages between aboveground and belowground biota[J]. Science，2004，304（5677）：1629-1633
[47] BAI Y F，COTRUFO M F. Grassland soil carbon sequestration： current understanding， challenges， and solutions[J]. Science，2022，377（6606）：603-608
[48] LAL R. Soil carbon sequestration impacts on global climate change and food security[J]. Science，2004，304（5677）：1623-1627
[49] LI Q，SONG Y T，ZHOU D W，et al. Effects of fencing and grazing on soil carbon， nitrogen， phosphorus storage in degraded alkali-saline grassland[J]. Pratacultural Science，2014，31（10）：1811-1819 李强，宋彦涛，周道玮，等.围封和放牧对退化盐碱草地土壤碳、氮、磷储量的影响[J].草业科学，2014，31（10）：1811-1819
[50] EVANS J R. Photosynthesis and nitrogen relationships in leaves of C3 plants[J]. Oecologia，1989，78（1）：9-19
[51] FORNARA D A，TILMAN D. Plant functional composition influences rates of soil carbon and nitrogen accumulation[J]. Journal of Ecology，2008，96（2）：314-322
[52] GENG X Q，TANG Y K，WANG L N，et al. Nitrogen addition increases biomass but reduces nitrogen use efficiency of terrestrial plants in China[J]. Chinese Journal of Plant Ecology，2024，48（2）：147-157 耿雪琪，唐亚坤，王丽娜，等.氮添加增加中国陆生植物生物量并降低其氮利用效率[J].植物生态学报，2024，48（2）：147-157
[53] SUN Y F，WANG Y P，YAN Z B，et al. Above-and belowground biomass allocation and its regulation by plant density in six common grassland species in China[J]. Journal of Plant Research，2022，135（1）：41-53
[54] KRAMER-WALTER K R，LAUGHLIN D C. Root nutrient concentration and biomass allocation are more plastic than morphological traits in response to nutrient limitation[J]. Plant and Soil，2017，416（1）：539-550
[55] ZHANG X，ZHOU Y Z，JI Y H，et al. Climate factors affect above-belowground biomass allocation in broad-leaved and coniferous forests by regulating soil nutrients[J]. Plants，2023，12（23）：3926
[56] SONG W J，SU J S，ZHANG M D，et al. Plant compensatory growth and optimal grazing intensity of grasslands in northern China： a meta-analysis of grazing experiments[J]. Chinese Science Bulletin，2023，68（11）：1330-1342 宋伟江，苏纪帅，张梦迪，等.中国北方草地植物补偿性生长与合理放牧强度：基于放牧实验的整合分析[J].科学通报，2023，68（11）：1330-1342
[57] WANG L，LUAN L M，HOU F J，et al. Nexus of grazing management with plant and soil properties in northern China grasslands[J]. Scientific Data，2020，7：39
[58] ZHANG Y J，ZHU J T，SHEN R N，et al. Research progress on the effects of grazing on grassland ecosystem[J]. Chinese Journal of Plant Ecology，2020，44（5）：553-564 张扬建，朱军涛，沈若楠，等.放牧对草地生态系统影响的研究进展[J].植物生态学报，2020，44（5）：553-564
[59] JIN M M，XU Z R，CHENG S K. Response of vegetation and soil to different grazing intensities of alpine grassland in Northern Tibet[J]. Acta Ecologica Sinica，2020，40（23）：8753-8762 靳茗茗，徐增让，成升魁.藏北高寒草地植被和土壤对不同放牧强度的响应[J].生态学报，2020，40（23）：8753-8762
[60] ZHANG C X，NAN Z B. Research progress on effects of grazing on physical and chemical characteristics of grassland soil[J]. Acta Prataculturae Sinica，2010，19（4）：204-211 张成霞，南志标.放牧对草地土壤理化特性影响的研究进展[J].草业学报，2010，19（4）：204-211
[61] ERDEN OCHIR. Study of soil physical properties and water characteristics change under different grazing intensities[D]. Hohhot：Inner Mongolia Agricultural University，2012：9-21 额尔登傲其尔.不同放牧强度下土壤物理性状及水分特征变化研究[D].呼和浩特：内蒙古农业大学，2012：9-21
[62] LI Y Q，ZHAO J B. Effects of overgrazing on ecological and environmental construction and measurement[J]. Journal of Desert Research，2005，（3）：404-408 李瑜琴，赵景波.过度放牧对生态环境的影响与控制对策[J].中国沙漠，2005，（3）：404-408
[63] DU Z Y，CAI Y J，WANG X D，et al. Research progress on grazing livestock dung decomposition and its influence on the dynamics of grassland soil nutrients[J]. Acta Ecologica Sinica，2019，39（13）：4627-4637 杜子银，蔡延江，王小丹，等.放牧牲畜粪便降解及其对草地土壤养分动态的影响研究进展[J].生态学报，2019，39（13）：4627-4637

高寒草甸不同放牧强度下植物地上-地下生物量权衡的变化

Changes of Trade-offs Between Aboveground and Belowground Plant Biomass Across Different Grazing Intensities of Alpine Meadow

PDF

可视化

摘要/Abstract

引用本文

使用本文

参考文献

相关文章 15

编辑推荐

Metrics

本文评价

[1]	段俊光, 房凯, 褚建民, 王迎新, 张琦. 退化程度对典型草原物种多样性和土壤养分关系的影响[J]. 草地学报, 2025, 33(9): 2890-2899.
[2]	郭芳, 张卫青, 关海波, 万志强, 袁亚楠, TSEDEVDORJ Ser-od, 张丽. 不同放牧强度下内蒙古典型草原地表节肢动物群落特征[J]. 草地学报, 2025, 33(9): 2950-2961.
[3]	赵海东, 梁海红, 胡小文, 冯彦皓, 常生华, 李春杰, 侯扶江. 基于草地植被恢复指数的退化高寒草甸生态修复措施初步效果评价[J]. 草地学报, 2025, 33(9): 3057-3067.
[4]	王瑞, 苏世锋, 尹亚丽, 赵文, 王彦龙, 李世雄. 放牧对高寒草甸春季草场土壤磷素转化率的影响[J]. 草地学报, 2025, 33(8): 2502-2511.
[5]	张丽, 张卫青, 关海波, 袁亚楠, 春风, 万志强, 郭芳. 不同放牧强度下内蒙古典型草原土壤团聚体线虫群落特征[J]. 草地学报, 2025, 33(8): 2531-2540.
[6]	张磊, 张玉娟, 孙世贤, 徐学宝, 邢越, 刘晓丽, 高翠萍, 王成杰. 不同放牧强度对荒漠草原土壤活性有机碳的影响[J]. 草地学报, 2025, 33(8): 2541-2547.
[7]	姜成栋, 李国荣, 童生春, 李进芳, 朱海丽, 刘亚斌, 陈文婷, 胡夏嵩. 黄河源高寒草甸鼠丘斑块土壤特性对植被多样性的影响[J]. 草地学报, 2025, 33(8): 2652-2665.
[8]	何升然, 刘晓静, 韩天虎, 赵雅姣, 韩颜隆. 紫花苜蓿氮效率差异对其根际土壤养分特性及微生物群落特征的影响[J]. 草地学报, 2025, 33(7): 2150-2161.
[9]	周慧敏, 高志伟, 张晓嘉, 吴佳芯, 王伟峰, 韩国栋. 不同放牧强度对内蒙古荒漠草原土壤细菌群落的影响[J]. 草地学报, 2025, 33(6): 1764-1770.
[10]	赵程鹏, 解婷婷, 单立山, 王海霞, 肖路臣, 何彩, 张斌. 降水和UV-B辐射对凋落物层下土壤养分及化学计量特征的影响[J]. 草地学报, 2025, 33(6): 1894-1902.
[11]	李艳, 徐靖杭, 吴建平. 生物炭添加对云南昭通退化人工草地生物量的影响[J]. 草地学报, 2025, 33(6): 1991-2001.
[12]	吕亮雨, 施建军, 蔡宗程, 保善存, 高佩, 李发毅, 付守全, 刘永璐. 粪肥配施化肥对退化高寒草原牧草生产和土壤养分的影响[J]. 草地学报, 2025, 33(6): 2023-2032.
[13]	王佳婧, 贺思腾, 梁新然, 杨梦娇, 张亚, 杨志新, 何永美. 大狼毒入侵对滇西北亚高山草甸土壤-植物碳氮磷生态化学计量的影响[J]. 草地学报, 2025, 33(5): 1523-1533.
[14]	戢爽, 吴昊坤, 张绍雄, 谢惠春. 不同海拔高寒草甸土壤细菌多样性特征及影响因素研究[J]. 草地学报, 2025, 33(4): 1085-1094.
[15]	王文碧, 刘梦瑶, 欧克杰, 郝彪彪, 龚俊强, 苏军虎. 高原鼢鼠（Eospalax baileyi）鼠丘土壤化学计量比和酶活性动态变化[J]. 草地学报, 2025, 33(4): 1127-1136.