Research on Niche， Interspecific Association and Community Stability of the Shrub-Herb Layer in the Wetlands of the Lhasa River Basin

doi:10.11733/j.issn.1007-0435.2026.04.020

Acta Agrestia Sinica ›› 2026, Vol. 34 ›› Issue (4): 1363-1375.DOI: 10.11733/j.issn.1007-0435.2026.04.020

Research on Niche， Interspecific Association and Community Stability of the Shrub-Herb Layer in the Wetlands of the Lhasa River Basin

FU Ben-yi^1,2,3,4, SONG Guo-fu⁵, REN Xiao-long⁶, SUN Xue-ting^1,2,3,4, LUO Da-qing^1,2,3,4, QU Xing-le^1,2,3,4

1. Institute of Plateau Ecology, Xizang Agricultural and Animal Husbandry University, Nyingchi, Xizang 860000, China;
2. Key Laboratory of Alpine Vegetation Ecological Security of Husbandry University, Nyingchi, Xizang 860000, China;
3. Key Laboratory of Xizang Plateau Forest Ecology of Ministry of Education & National, Nyingchi, Xizang 860000, China;
4. Ngari Ecological Monitoring Station, Ngari, Xizang 860000, China;
5. Xizang Autonomous Region Remote Sensing Monitoring Center for Ecological Environment, Lhasa, Xizang 850000, China;
6. Linzhi Municipal Ecology and Environment Bureau, Nyingchi, Xizang 860000, China

Received:2025-05-06 Revised:2025-06-18 Published:2026-04-15

拉萨河流域湿地灌草层生态位与种间联结及群落稳定性研究

付本艺^1,2,3,4, 宋国富⁵, 任晓龙⁶, 孙雪婷^1,2,3,4, 罗大庆^1,2,3,4, 屈兴乐^1,2,3,4

1. 西藏农牧大学高原生态研究所, 西藏林芝 860000;
2. 西藏自治区高寒植被生态安全重点实验室, 西藏林芝 860000;
3. 西藏高原森林生态教育部重点实验室, 西藏林芝 860000;
4. 阿里生态监测站, 西藏阿里 859000;
5. 西藏自治区生态环境遥感监测中心, 西藏拉萨 850000;
6. 林芝市生态环境局, 西藏林芝 860000

通讯作者: 屈兴乐，E-mail：1318266529@qq.com
作者简介:付本艺（1999-），男，汉族，山东济南人，主要从事群落生态学方面研究，E-mail：438524206@qq.com；
基金资助:
西藏农牧大学林学博士点（一期）（533325001）；第二次青藏高原综合科学考察研究项目（2019QZZK1006）；西藏农牧学院2025年学科建设项目（高原生态研究所生态学-YJSXK2025-01）；西藏农牧学院2025年学科建设项目（高原生态研究所资源利用与植物保护（资源利用方向）-YJSXK2025-16）资助

Abstract

Abstract: This study focused on the wetland ecosystem of the Black-necked Crane National Nature Reserve in the Lhasa River Basin， systematically explores the niche characteristics， interspecific associations， and community stability of the dominant populations in the shrub layer and herb layer， analyzes the survival strategies， resource competition， and symbiotic relationships of dominant species in the alpine wetland environment， aiming to promote the positive associations between dominant species and companion species and optimize the community structure； The results show that： the average value of the Levins index in the herb layer （7.97） is higher than that in the shrub layer （3.27）， and the average value of the Shannon index （2.03） is also higher than that in the shrub layer （1.20）， indicating that herbaceous plants have better breadth and uniformity of resource utilization. The niche overlap index shows that the average value in the herb layer （0.22） is higher than that in the shrub layer （0.18）， with more intense resource competition. The interspecific association analysis shows that the shrub-herb layer presents a positive association as a whole （all VR values are greater than 1）. The overall symbiosis of plants in the shrub layer is stronger than competition， while the resource competition among species in the herb layer is more intense than that in the shrub layer. The community stability model shows that the Euclidean distances between the shrub layer， the herb layer and the theoretical stable point （20， 80） are 7.98 and 22.77 respectively， and the herb layer is in an unstable state. The study suggests optimizing vegetation configuration by screening species with matching ecological habits to enhance community stability， so as to provide a theoretical basis for the vegetation management of the reserve and the protection of the Qinghai-Xizang Plateau ecosystem.

Key words: Lhasa River Basin of wetlands, Shrub-herb layer, Niche characteristics, Interspecific association, Community stability

摘要： 本研究聚焦拉萨河流域黑颈鹤国家自然保护区湿地生态系统，对灌木层和草本层优势种群的生态位特征、种间联结及群落稳定性展开系统探究，分析高寒湿地环境下优势物种的生存策略、资源竞争与共生关系，以促进优势种与伴生种的正联结，优化群落结构。研究结果表明：草本层 Levins 指数均值（7.97）高于灌木层（3.27），Shannon 指数均值（2.03）亦高于灌木层（1.20），表明草本植物资源利用广度与均匀性更优；生态位重叠指数显示，草本层均值（0.22）高于灌木层（0.18），资源竞争更为激烈；种间联结分析表明，灌草层整体呈正联结（VR均大于1），灌木层植物整体共生大于竞争，草本层物种对于资源竞争较灌木层激烈。群落稳定性模型显示，灌草层与理论稳定点（20，80）的欧式距离分别为 7.98 和 22.77，草本层处于不稳定状态。研究建议通过筛选生态习性匹配物种优化植被配置，提升群落稳定性，为保护区植被管理、高原生态系统保护提供理论依据。

关键词: 拉萨河流域湿地, 灌草层, 生态位特征, 种间联结, 群落稳定性

CLC Number:

S812

FU Ben-yi, SONG Guo-fu, REN Xiao-long, SUN Xue-ting, LUO Da-qing, QU Xing-le. Research on Niche， Interspecific Association and Community Stability of the Shrub-Herb Layer in the Wetlands of the Lhasa River Basin[J]. Acta Agrestia Sinica, 2026, 34(4): 1363-1375.

付本艺, 宋国富, 任晓龙, 孙雪婷, 罗大庆, 屈兴乐. 拉萨河流域湿地灌草层生态位与种间联结及群落稳定性研究[J]. 草地学报, 2026, 34(4): 1363-1375.

References

[1] YIN Z H，CHEN XF，LA D. Analysis on the niche and interspecific association of dominant plant species in alpine meadow under simulated warming and grazing[J]. Acta Agrestia Sinica，2023，31（5）：1302-1313 尹正辉，陈新丰，拉多. 模拟增温和放牧对高寒草甸主要植物生态位及种间联结的影响[J]. 草地学报，2023，31（5）：1302-1313
[2] ZHAN Y F，MA L，LI X Y，et al. Niches of dominant populations of wetland plants in Zhangye section of Heihe Rive[J]. Journal of Northeast Forestry University，2012，40（10）：43-47 占玉芳，马力，李小燕，等. 黑河流域（张掖段）湿地植物群落优势种群生态位[J]. 东北林业大学学报，2015，43（1）：43-47
[3] LIU J M，LIU H，LI L X. Resource utilization characteristics of dominant species in Cinnamomum migao community[J]. Bulletin of Botanical Research，2021，41（3）：380-386 刘济明，刘欢，李丽霞.米槁群落主要物种资源利用特征分析[J]. 植物研究，2021，41 （3）： 380-386
[4] YU Y，BAI X J，WANG Z Y. Niche characteristics and competitive relationship of Larix gmelinii between different age classes in Greater Khingan Range secondary forest area[J]. Acta Ecologica Sinica，2022，42（12）：4912-4921 俞昀，白小军，王志一.大兴安岭次生林区不同龄级落叶松（Larix gmelinii）生态位特征和竞争关系[J]. 生态学报，2022，42（12）： 4912-4921
[5] LI Z H， ZHAO Y Y， WU S， et al. Analysis on niche and interspecific association of dominant species in deciduous broad-leaved forest of Beijing Sizuolou Municipal-level nature reserve[J]. Journal of Plant Resources and Environment， 2025， 34（4）： 83-94 李子涵，赵悦悦，吴爽，等. 北京四座楼市级自然保护区落叶阔叶林优势种生态位及种间联结分析[J]. 植物资源与环境学报，2025，34（4）：83-94
[6] YAN X H，TUO H H，LI M H，et al. Interspecific association of plants in the process of the subshrub expansion in typical steppe of Yunwu Mountain，Ningxia[J]. Acta Agrestia Sinica，2023，31（1）：230-238 晏昕辉，拓行行，李美慧，等. 宁夏云雾山典型草原半灌木扩张过程中植物种间联结性研究[J].草地学报， 2023， 31（1）： 230-238
[7] QI X Y，WANG X Y，YIN Z M，et al. Relationship and mechanism between species diversity and community stability in long-term grassland[J]. Acta Agrestia Sinica，2025，33（4）：1239-1248 祁旭阳，王欣雨，尹子鸣，等. 长时间尺度草地物种多样性-群落稳定性关系及机制[J]. 草地学报， 2025， 33（4）： 1239-1248
[8] LI D Y， LÜ B L， JIANG Y， et al. Influence of tending thinning on niche characteristics and interspecific association of planted forest vegetation in saihanba nature reserve[J]. Journal of Arid Land Resources and Environment， 2025， 39（1）： 139-149 李丹阳，吕宝磊，姜雨，等. 抚育间伐对塞罕坝自然保护区人工林植被生态位特征和种间关联性的影响[J].干旱区资源与环境，2025，39（1）：139-149
[9] CAI J Y，ZHENG D G，LUO G，et al. Nicheand community stability of dominant species in the tree-shrub-grass layer of Quercus aliena var. acutiserrata community[J]. Journal of Hunan Ecological Science，2023，10（1）：44-54 蔡京勇，郑德国，罗刚，等. 锐齿槲栎群落乔灌草层优势种生态位及群落稳定性[J]. 湖南生态科学学报，2023，10 （4）： 34-42
[10] WANG S， ISBELL F， DENG W， et al. E. How complementarity and selection affect the relationship between ecosystem functioning and stability[J]. Ecology， 2021，102（12）： e03347
[11] GODOY O， GÓMEZ-APARICIO L， MATÍAS L. An excess of niche differences maximizes ecosystem functioning[J]. Nature Communications， 2020，11：4312
[12] CAO S X， HUANG Y T， LUO Z， et al. Analysis on niche characteristics of dodonaea viscosa shrub communities in fujian islands and environmental impacts on them[J]. Journal of Northeast Forestry University， 2025， 53（4）： 88-98 曹晟烜，黄运腾，罗真，等. 福建海岛车桑子灌丛群落生态位特征及环境对其影响[J]. 东北林业大学学报，2025，53（4）：88-98
[13] JIANG H X， GUO D C， LIU M H， et al. Study on diversity and niche of macrozoobenthos in Zhongyangzhan Black-billed capercaillie national nature reserve[J]. Wetland Science， 2024， 22（4）： 508-517 姜海霞，郭东昌，刘曼红，等. 中央站黑嘴松鸡国家级自然保护区大型底栖动物多样性和生态位研究[J]. 湿地科学，2024，22（4）：508-517
[14] LIU M J， LI H J. Rare wildlife resources and their protection in the black-necked crane national nature reserve in the middle reaches valley of the Yarlung Zangbo River， Xizang[J]. Journal of Central South Forest Inventory and Planning， 2020， 39（2）： 57-61 刘敏杰，李华军. 西藏雅鲁藏布江中游河谷黑颈鹤国家级自然保护区珍稀野生动植物资源与保护[J]. 中南林业调查规划，2020，39（2）：57-61
[15] WANG T，SHEN W S. The temporal and spatial changes of temperature in Black-necked Crane over wintering area of Yarlung Zangbo River[J]. Acta Ecologica Sinica，2020，40（11）：3560-3570 王涛，沈渭寿.雅鲁藏布江黑颈鹤越冬地气温时空变化特征[J]. 生态学报，2020，40（11）：3560-3570
[16] ZHANG J T. Quantitative ecology[M]. 3rd Edition. Beijing： Science Press， 2018： 13-216 张金屯. 数量生态学[M].第3 版.北京：科学出版社，2018：13-216
[17] LEVINS R. Evolution in changing environments： some theoretical explorations[M]. Princeton： Princeton University Press，1968：32-120
[18] SILVERTOWN J，DODD M， GOWING D. Hydrological niches in terrestrial plant communities： A review[J]. Journal of Ecology，2015，103（1）：93-108
[19] SCHOENER T W. Resource partitioning in ecological communities： research on how similar species divide resources helps reveal the natural regulation of species diversity [J]. Science， 1974，185（4145）： 27-39
[20] PIANKA E R. The structure of lizard communities [J]. Annu Rev Ecol Syst，1973，4（1）： 53-74
[21] SCHLUTER D. A variance test for detecting species association，with some example applications.Ecology，1984，65（3）： 998-1005
[22] ALAM， YIN Z H， LI S， et al. Analysis on niche and interspecific association of dominant herbs in swamp wetlands of the Lhasa River Basin[J]. Acta Agrestia Sinica， 2025， 33（8）： 2666-2676 阿拉姆，尹正辉，李森，等. 拉萨河流域沼泽湿地优势草本生态位及种间联结分析[J]. 草地学报，2025，33（8）：2666-2676
[23] CHEN L H， ZHAO Z G， ZHOU M. Niche and interspecific association of herbaceous plant communities in the outer Qinhuai River of Nanjing[J]. Journal of Nanjing Forestry University （Natural Sciences Edition）， 2023， 47（6）： 203-211 陈丽华，赵志刚，周敏. 南京外秦淮河草本植物群落生态位与种间联结性[J].南京林业大学学报（自然科学版），2023，47 （6）：203-211
[24] ZHENG Y R. Comparison of methods for studying stability of forest community[J]. Scientia Silvae Sinicae，2000，36（5）：28-32 郑元润.森林群落稳定性研究方法初探[J].林业科学，2000，36（5）： 28-32
[25] DU Z Y， XING W L， XUE L， et al. Niche characteristics of main species in plant communities and their interspecific association in antimony mining areas with karst rocky desertification[J]. Acta Ecologica Sinica， 2023， 43（7）： 2865-2880 杜忠毓，邢文黎，薛亮，等. 喀斯特石漠化锑矿区植物群落主要物种生态位特征及其种间联结[J]. 生态学报，2023， 43 （7）： 2865-2880
[26] LI Z J. Community ecology[M]. Beijing： China Meteorological Press， 2001：李振基. 群落生态学[M].北京：气象出版社，2001：
[27] SONG H Y， SUN C L， CHAI Z Z， et al. Niche and interspecific relationships of dominant herbaceous plant populations in lead-zinc mine waste dumps in Northwest Guizhou[J]. Acta Agrestia Sinica， 2022， 30（10）： 2764-2771 宋红艳，孙彩丽，柴宗政. 黔西北铅锌矿废渣场优势草本植物种群生态位及种间关系[J]. 草地学报， 2022， 30（10）： 2764-2771
[28] SHI H B，CHEN Y N，LI W H，et al. Interspecific association and stability of vegetation in the lower reaches of the Tarim River[J]. Arid Zone Research，2020，37（1）：220-226 史浩伯，陈亚宁，李卫红，等. 塔里木河下游植被种间关系与稳定性分析[J]. 干旱区研究，2020，37（1）： 220-226
[29] SUN J， ZHU X D， CAI X C， et al. Short-term effects of fertilization and reseeding on plant community stability and interspecific relationships in alpine meadows[J]. Acta Agrestia Sinica， 2026，34（1）：275-286 孙舰，朱熙栋，蔡新成，等. 施肥补播对高寒草甸植物群落稳定性及种间关系的短期影响[J]. 草地学报，2026，34（1）：275-286
[30] LU Y， WU W P， ZENG Y. Niche characteristics and interspecific association of dominant plants in the oasis-desert ecotone on the northern foot of the Tianshan Mountains[J]. Journal of Desert Research， 2024， 44（2）： 254-3263 卢燕，吴万平，曾勇. 天山北麓绿洲荒漠过渡带优势植物的生态位特征与种间联结[J]. 中国沙漠，2024， 44（2）：254-3263
[31] LEI D F，WANG Z J. Interspecific associations，niches and community stability of main species in Pinus massoniana community in karst mountainous cities，the central Guizhou Province[J]. Acta Ecologica Sinica，2024，44（10）：4353-4367 雷德芳，王志杰. 黔中喀斯特地区马尾松群落主要物种种间联结、生态位及群落稳定性特征[J].生态学报，2021，41（14）：5521-5530
[32] FANG J P，LU J，LUO J，et al. Study on the ecological niche of psychrophyte Rhodiola fasitiata population[J]. Journal of Anhui Agricultural Sciences，2009，37（5）：2027-2031 方江平，卢杰，罗建，等. 高寒植物长鞭红景天种群生态位研究[J]. 西藏农业科技，2009，31 （5）： 2027-2031
[33] BAI Y， WU J， CLARK C M， et al. Ecosystem stability and compensatory effects in the Inner Mongolia grassland[J]. Nature， 2004， 431（7005）： 181-184
[34] GAO L， JIANG Z H， HONG Z C， et al. Analysis on niche and interspecific association of main tree species in castanopsis eyrei forest of Wuyi Mountain National Park[J]. Journal of Plant Resources and Environment， 2024， 33（6）：65-73 高亮，江子涵，洪子辰，等. 武夷山国家公园甜槠林主要树种生态位与种间联结分析[J]. 植物资源与环境学报，2024，33（6）：65-73
[35] GAO Y H，ZENG X Y，ZHOU G Y，et al. Interspecific relationships of dominant populations in the alpine wetlands in the source region of Yangtze River[J]. Wetland Science，2011，9（1）：1-7 高永恒，曾晓阳，周国英，等. 长江源区高寒湿地植物群落主要种群种间关系分析[J]. 湿地科学，2011，9 （1）： 1-7
[36] XIONG M， AN H B， ZHAO M L， et al. Influence of grazing on spatial pattern and niche of main plant populations in desert steppe[J]. Acta Agrestia Sinica， 2024， 32（4）： 1177-1183 熊梅，安海波，赵萌莉，等. 放牧对荒漠草原主要植物种群空间格局与生态位的影响[J]. 草地学报， 2024， 32（4）： 1177-1183
[37] WANG J J， ZHU J H， GAO X， et al. Analysis on plant species diversity and interspecific association of typical wetlands in the middle and lower reaches of the Lhasa River[J]. Acta Agrestia Sinica， 2024， 32（3）： 918-927 王净杰，朱建航，高鑫，等. 拉萨河中下游典型湿地植物物种多样性及种间联结性分析[J]. 草地学报， 2024， 32（3）： 918-927
[38] HE J， XU L X， AI X R， et al. Dynamics of niche and interspecific association of dominant species in natural secondary forest of Xingdou Mountain[J]. Journal of Forestry Research， 2024， 37（1）： 171-182 何江，徐来仙，艾训儒，等. 星斗山天然次生林优势种的生态位和种间联结动态[J]. 林业科学研究，2024，37 （1）： 171-182
[39] LIU Y T，HOU M F，HE L Y，et al. Niche and interspecific association of dominant tree species in karst forest of Junzi Mountain，Eastern Yunnan，China[J]. Chinese Journal of Applied Ecology，2023，34（7）：1771-1778 刘雨婷，侯满福，贺露炎，等.滇东菌子山喀斯特森林群落乔木优势树种生态位和种间联结[J].应用生态学报，2023，34（7）： 1771-1778
[40] FARGIONE J E，TILMAN D. Diversity decreases invasion via both sampling and complementarity effects[J]. Ecology Letters，2005，8（6）：604-611
[41] MACDOUGALL A S， GILBERT B， LEVINE J M. Plant invasion and the niche[J]. Journal of Ecology， 2009，97（4）： 609-615
[42] WU M J， QIU J， ZHENG F， et al. Study on shrub species diversity and niche of Xinjiang Wild Fruit Forest[J]. Arid Zone Research， 2024， 41（12）： 2094-2109 吴明江，邱娟，郑凤，等. 新疆野果林灌木物种多样性与生态位研究[J]. 干旱区研究，2024，41（12）：2094-2109
[43] ZHANG Z H，HU G. Interspecific relationship analysis of dominant species in Quercus Cyclobalanopsis community in karst mountain area[J]. Ecology and Environmental Sciences，2011，20（S2）：1209-1213 张忠华，胡刚. 喀斯特山地青冈栎群落优势种的种间关系分析[J].生态环境学报，2011，20（S2）： 1209-1213
[44] HU Y， WANG H.C，JIA H，et al. Ecological niche and interspecific association of plant communities in alpine desertification grasslands： A case study of Qinghai Lake Basin[J]. Plants （Basel）， 2022，11（20）：2724
[45] HU N N， CUI Y X， SUN M， et al. Niche and interspecific association of dominant populations in the shrub layer of typical forest communities in the western section of the Qinling Mountains[J]. Chinese Journal of Ecology， 2024， 43（12）： 3595-3604 胡宁宁，崔雨昕，孙萌，等. 秦岭西段典型森林群落灌木层优势种群生态位与种间联结[J]. 生态学杂志，2024， 43 （12）： 3595-3604
[46] ZHOU H K，ZHOU L，ZHAO X Q，et al. Study on the stability of alpine meadow ecosystem in Qinghai-Tibet Plateau[J]. Chinese Science Bulletin，2006，51（1）：63-69 周华坤，周立，赵新全，等.青藏高原高寒草甸生态系统稳定性研究[J].科学通报，2006，51（1）：63-69

Research on Niche， Interspecific Association and Community Stability of the Shrub-Herb Layer in the Wetlands of the Lhasa River Basin

拉萨河流域湿地灌草层生态位与种间联结及群落稳定性研究

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